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106 Farsad Nadjafi and Mohammad Reza Kanani
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foenugraeca (L.) E.H.L. Krause, Telis foe­num-graecum (L.) Kuntze, Trigonella ensifera
Trautv., T. graeca St.-Lag., T. haussknechtii (Širj.) Vassilcz., T. jemenensis (Serp.) Sin­skaya, T. rhodantha (Alef.) Vassilcz., T. tibet- ana (Alef.) Vassilcz., Xiphostylis erectus Gasp.
Family: Fabaceae (Leguminosae).
Arabic name: Holbah.
Persian name: Shanbalileh.
English name: Fenugreek (Fig. 2.72).
Distribution
Africa: Algeria, Egypt, Ethiopia, Kenya, Libya, Mali, Morocco, Mozambique, Sudan, Tanza­nia, Tunisia, Zimbabwe. Asia: Afghanistan, Austria, Bangladesh, Caucasus, China, India, Iran, Iraq, Kuwait, Kyrgyzstan, Lebanon, Mongolia, Myanmar, Nepal, Oman, Pakistan, Saudi Arabia, Syria, Tajikistan, Transcauca­sus, Turkey, Turkmenistan, Uzbekistan, Yemen. Europe: Albania, Baltic States, Bel­gium, Bulgaria, Cyprus, France, Germany,
Greece (Aegean Islands), Hungary, Italy, Por­tugal, Romania, Russia, Spain, Switzerland, Ukraine. Oceania and Australasia: Fiji.
Morphology
Annual herb, 30–80 cm. Taproot to 80 cm, fibrous roots lateral. Stem erect, terete, branched. Leaves 3-foliolate; stipules entire, membranous, base adnate to petiole, apex acute; petiole spreading, 6–15 mm; leaflets narrowly obovate, ovate, to oblong-elliptic, terminal equal to laterals, 15–40 mm × 4–15 mm, sparsely pubescent or glabrescent abax­ially, glabrous adaxially, lateral veins 5 or 6 pairs, base cuneate, margin dentate in upper half. Flowers sessile, axillary, solitary or in pairs. Calyx 7–8 mm, villous. Corolla creamy or pale yellow, base violet, 13–18 mm. Ovary puberulent; ovules numerous. Legume long conical, 7–12 cm × 0.4–0.5 cm, straight or slightly curved, glabrescent, veins longitu­dinally reticulate, apex with long beak to 2 cm, distal suture thickened. Seeds 10–20, brown, oblong-ovoid, 3–5 mm × 2–3 mm, surface uneven (Bobrov et al., 1934–1964; Rechinger, 1963–2001; Tutin et al., 1964; Davis, 1965–1988).
Fig. 2.72. Trigonella foenum-graceum L. (By Yak, CC BY-SA 3.0, https://de.wikipedia.org/wiki/ Bockshornklee#/media/Datei:Junger_ Bockshornklee.jpg.)
Urtica urens L., Sp. Pl.: 984 (1753)
Synonyms: Urtica atlantica Blume, U. cuben- sis Klotzsch ex Herder, U. minor Garsault, U. minor Lam., U. monoica Gilib., U. ovalifolia
Stokes, U. parvula Blume, U. quadristipulata Dulac, U. trianae Rusby.
Family: Urticaceae.
Arabic name: Qoras hareq.
Persian name: Gazanahe sag.
English name: Annual nettle, Nettle, Dog nettle (Fig. 2.73).
Distribution
Africa: Algeria, Egypt, Eritrea, Ethiopia, Kenya, Lesotho, Libya, Morocco, Namibia,
Botany, Collection, and Cultivation 107
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elliptic, sometimes ovate or obovate, 1.2–6
cm × 0.6–3 cm, 5-veined, often subglabrous
except for sparse stinging hairs on both sur-
faces, base broadly cuneate or rounded, mar-
gin 6–11-dentate, apex obtuse-rounded;
cystoliths punctiform, distinct adaxially. In-
florescences containing proximal female
flowers and distal male flowers, spicate,
0.5–2.5 cm. Male flowers short pedicellate,
in bud c.1.2 mm; perianth lobes connate half
of length, puberulent. Female flowers: peri
anth lobes connate at base, dorsal-ventral
lobes ovate, equaling achene, often with 1
stinging hair on dorsal rib, sparsely setulose
along margin, lateral lobes ovate, c.5× as
small as others. Achene brownish-gray,
ovoid, compressed, 0.8 mm, verrucose, in-
vested by persistent perianth lobes (Bobrov
et al., 1934–1964; Rechinger, 1963–2001;
Tutin et al., 1964; Davis, 1965–1988; Sell
and Murrell, 2006).
-
3.0, https://upload.wikimedia.org/wikipedia/ commons/6/61/Urtica_urens_002.JPG.)
Somalia, Sudan, Tanzania, Tunisia. Asia: China , Caucasus, Bahrain, Kuwait, Iran, Iraq, Kaz­akhstan, Kyrgyzstan, Lebanon, Oman, Qatar, Saudi Arabia, Syria, Transcaucasus, Turkey, Yemen. Europe: Albania, Austria, Baltic States, Belarus, Belgium, Bulgaria, Canary Islands, Great Britain, Cyprus, Denmark
Greece (Aegean Islands), Hungary, Ireland, Italy, Netherlands, Norway, Poland, Portu­gal, Romania, Russia, Spain, Sweden, Switz­erland, Ukraine. Americas: Argentina, Bolivia, Brazil, Canada, Chile, Colombia, Cuba, Ecua­dor, Guatemala, Mexico, Peru, USA. Oceania and Australasia: Australia, New Zealand.
Morphology
Annual herb. Stems branched, 10–60 cm tall, sparsely puberulent and somewhat densely armed with stinging hairs; lower internodes 4–7 cm, upper internodes 1–3 cm. Stipules free, narrowly triangular, 1–2.5 mm, ciliate; petiole 1–2.5 cm, puberulent, sparsely armed with stinging hairs; leaf blade broadly
Valeriana officinalis L., Sp. Pl.: 31 (1753)
Synonyms: Astrephia chinensis Dufr., Valeria-
na alternifolia Bunge, V. alternifolia f. angusti-
folia (Kom.) Kitag., V. alternifolia f. verticillata
(Kom.) S.X. Li, V. alternifolia subsp. pseu-
docoreana Vorosch., V. alternifolia subsp.
stubendorfiana (Sumn.) Vorosch., V. alternifo-
lia subsp. stubendorfii (Kreyer ex Kom.) Vor-
osch., V. alternifolia var. angustifolia (Kom.)
S.H. Li, V. alternifolia var. stolonifera A.I. Bar-
anov & Skvortsov, V. altissima J.C. Mikan, V.
altissima J.C. Mikan ex Besser, V. angustifolia
Tausch ex Host, V. baltica Pleijel, V. chinensis
Kreyer, V. chinensis Kreyer ex Kom., V. coreana
Briq., V. coreana subsp. badzhalensis Vorosch.,
V. coreana subsp. leiocarpa (Kitag.) Vorosch.,
V. coreana subsp. pseudoumbrosa (Vorosch.)
Vorosch., V. coreana subsp. sachalinensis
(Hara) Vorosch., V. divaricata Huter, V. exal-
tata J.C. Mikan, V. exaltata J.C. Mikan ex
Pohl, V. fauriei Briq., V. fauriei f. coreana
(Briq.) H. Hara, V. fauriei subsp. badzhalensis
(Vorosch.) Vorosch., V. fauriei subsp. sa-
chalinensis (Hara) Vorosch., V. fauriei var.
dasycarpa H. Hara, V. fauriei var. fauriei,
V. fauriei var. leiocarpa (Kitag.) Kitag., V.
jacutica Sumn., V. leiocarpa Kitag., V. lucida
108 Farsad Nadjafi and Mohammad Reza Kanani
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Hort. Par., V. lucida Hort. Par. ex DC., V. major Pall., V. multiceps Wallr., V. multiceps
var. dentata Wallr., V. multiceps var. incisa Wallr., V. nipponica Nakai, V. nipponica Nakai ex Kitag., V. officinalis f. pilosa Podp., V. offic-
inalis f. pubescens Podp., V. officinalis subsp. baltica (Pleijel) Á. Löve & D. Löve, V. officina- lis subsp. communis Schübl. & G. Martens, V. officinalis subsp. dunensis P.D. Sell, V. offic- inalis subsp. exaltata (J.C. Mikan ex Pohl)
Wirtg., V. officinalis subsp. exaltata (J.C. Mi- kan) Soó, V. officinalis subsp. latifolia (Vahl) Schübl. & G. Martens, V. officinalis subsp. sarkayi (Soó) Á. Löve & D. Löve, V. officinalis subsp. simplicifolia Ledeb., V. officinalis sub- sp. vulgaris Celak., V. officinalis var. alternifo- lia Ledeb., V. officinalis var. altissima (J. C . Mikan ex Besser) W.D.J. Koch, V. officinalis var. angustifolia Hayne, V. officinalis var. an- gustifolia Miq., V. officinalis var. exaltata (J. C. Mikan ex Pohl) Kostel., V. officinalis var. lat-
ifolia Briq., V. officinalis var. latifolia Miq., V. officinalis var. latifolia Vahl, V. officinalis var. major Wimm. & Grab., V. officinalis var. media W.D.J. Koch, V. officinalis var. officina- lis, V. officinalis var. palustris Wibel, V. offici­nalis var. sarkanyi Soó, V. officinalis var. sarkayi Soó, V. officinalis var. simplicifolia Ledeb., V. officinalis var. vulgaris Kitt., V. palustris Kreyer, V. palustris var. exaltata (J.C. Mikan ex Pohl) Kreyer, V. palustris var. integella Kreyer, V. palustris var. vulgaris Kreyer, V. pinnata Gilib., V. provisa Kreyer, V. pseudofficinalis C.Y. Cheng & H.B. Chen, V. pseudoumbrosa Vorosch., V. pulchra Nakai, V. sambucifolia Eichw., V. sambucifolia subsp. sachalinensis Hara, V. sambucifolia var. dasy- carpa (Hara) H. Hara, V. sambucifolia var. fauriei (Briq.) H. Hara, V. sambucifolia var. sachalinensis Hara, V. sinensis J.F. Gmel., V. stubendorfiana Sumnev., V. stubendorfii Kreyer, V. stubendorfii Kreyer ex Kom., V. stubendorfii f. angustifolia Kom., V. stuben­dorfii f. verticillata Kom., V. stubendorfii sub- sp. angustifolia Kom., V. stubendorfii subsp. verticillata Kom., V. sylvestris Garsault, V. sylvestris Groschke, V. tergemina Kreyer, V. tianschanica Kreyer, V. tianschanica Kreyer ex Hand.-Mazz., V. vulgaris Bubani, Veronica officinalis var. altissima W.D.J. Koch.
Family: Caprifoliaceae.
Arabic name: Nārdin makhzani.
Persian name: Sonbolotib .
Common name: Valerian (Fig. 2.74).
Distribution
Asia: Caucasus, Iran, Transcaucasus, Turkey. Europe: Albania, Austria, Baltic States, Bela-
rus, Bulgaria, Denmark, Estonia, Finland, France, Germany, Greece, Hungary, Italy, Latvia, Lithuania, Poland, Romania, Russia, Sweden, Switzerland, Ukraine. Americas: USA.
Morphology
Plants up to 150 cm tall, infrequently sto­loniferous. Rhizomes reduced; roots fas­cicled, clavate, 1.5–2.5 mm in diameter. Stems hirsute or hispidulous, particularly at nodes, rarely glabrous. Basal and lowermost cauline leaves usually wilted at anthesis; cauline leaves petiolate to sessile; petiole 0–8 cm; blade ovate to broadly ovate, 5–15 cm
Fig. 2.74. Valeriana officinalis L. (By H. Zell, CC BY-SA 3.0, https://upload.wikimedia.org/wikipedia/ commons/8/82/Valeriana_officinalis_001.JPG.)
Botany, Collection, and Cultivation 109
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× 3–8 cm, pinnatisect; segments 7–15, lan­ceolate or linear, glabrous or ± hispidulous, base decurrent, margin entire or coarsely serrate; terminal and lateral segments simi­lar in shape and size. Inflorescence corymbi­form, large and compact even in fruit, or with 2 distinct, secondary lateral inflores­cences; bracts leaflike below, small and line­ar-lanceolate otherwise; bracteoles ellip­tic-oblong, oblanceolate, or linear-lanceolate, margin scarious-ciliate, apex awned. Corolla purplish-red, pink, or white, funnelform, 4–6 mm; lobes elliptic. Stamens and style strongly exserted. Achenes narrowly ovoid, 4–5 mm, both sides glabrous or pubescent (Boissier, 1867–1888; Bobrov etal., 1934– 1964; Rechinger, 1963–2001; Tutin et al., 1964; Davis, 1965–1988).
Veratrum album L., Sp. Pl.: 1044 (1753)
Synonyms: Helleborus albus (L.) Gueldenst., Melanthium album (L.) Thunb., M. bracteo­lare Desr., M. virens Thunb., Veratrum album albicans Gaudin, V. album f. glabrescens
(Zapal.) Soó, V. album subsp. misae (Sirj.) Tz- velev, V. album f. podolicum (Zapal.) Soó, V.
bosniacum Beck, V. croaticum (Beck) O. Loes., V. flavum (Griseb.) O. Loes., V. lobelianum f. oppositifolium Cheshm., V. misae (Sirj.) O.
Loes., V. parviflorum Bong., V. polygamum Gilib., V. viride Röhl.
Family: Melanthiaceae.
Arabic name: Khrbaq abyad.
Persian name: Khrbaqe sefid.
Common name: False helleborine, White hellebore, European white hellebore, White veratrum (Fig. 2.75).
Distribution
Asia: Transcaucasus, Turkey. Europe: Alba­nia, Austria, Baltic States, Belarus, Finland, France, Germany, Hungary, Italy, Norway, Poland, Portugal, Romania, Russia, Spain, Switzerland, Ukraine.
Fig. 2.75. Veratrum album L. (By Tigerente, CC
BY-SA 3.0, https://upload.wikimedia.org/wikipedia/
commons/8/80/Veratrum_album_Dachstein.jpg.)
Morphology
Stems 50–175 cm. Cauline leaves glabrous
or subglabrous above, shortly pubescent
beneath especially on the veins, the lower
10–25 cm × 6–15 cm, elliptical to broadly el-
liptical, becoming shorter and narrower
above. Inflorescence a much-branched pan-
icle. Pedicels 2–3 mm. Flowers 15–25 mm in
diameter. Perianth segments 7–15 mm ×
3–6 mm, elliptical to lanceolate, white out-
side and greenish inside to greenish-white
or yellowish on both surfaces. Capsule
sparsely pubescent (Bobrov et al., 1934–
1964; Tutin etal., 1964; Davis, 1965–1988).
Viola odorata L., Sp. Pl.: 934 (1753)
Synonyms: Viola cyclophylla Gand., V. g onz a -
loi Sennen, V. gonzaloi var. semiglabra Sen-
nen, V. hirta var. odorata (L.) Fiori, V. hirta
subsp. odorata (L.) Fiori, V. martia Gilib., V.
martia Hegetschw., V. martii subsp. odorata
(L.) Schimp. & Spenn., V. odorata f. odorata,
110 Farsad Nadjafi and Mohammad Reza Kanani
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V. odorata subsp. rierolae Sennen, V. odorata subsp. sarotina P. Fourn., V. parmensis Bailly, V. spectabilis K. Richt., V. tenerrima Halácsy & Braun, V. vilmoriniana Delacour & Mottet, V.
odorata proles brarcinonensis Sennen, V. od o ­rata subsp. texidoris Sennen, V. taynensis
Elisafenko.
Family: Violaceae.
Arabic name: Albanafsaji.
Persian name: Banafsheh.
English name: Sweet violet (Fig. 2.76).
Distribution
Africa: Algeria, Morocco, Tunisia. Asia: Caucasus, China, India, Iran, Iraq, Japan, Kazakhstan, Lebanon, Myanmar, Pakistan, Syria, Tajikistan, Transcaucasus, Turkey, Vietnam. Europe: Albania, Austria, Baltic States, Belarus, Belgium, Bulgaria, Canary Islands, Cyprus, Denmark, Finland, France, Germany, Great Britain, Greece (Aegean Is­lands), Hungary, Ireland, Italy, Nether­lands, Poland, Portugal, Romania, Russia, Spain, Sweden, Switzerland, Ukraine. Americas: Argentina, Canada, Chile, Co­lombia, Costa Rica, Cuba, Dominican Re­public, Ecuador, El Salvador, Guatemala, Leeward Islands, Mexico, Panamá, Peru, Puerto Rico, Trinidad and Tobago, USA, Venezuela.
Morphology
Perennial plant 5–15 cm, with leaf rosette and long, procumbent, rooting stolons. Leaves orbicular-reniform, deeply cordate, widest at about the middle; petiole long. Stipules ovate, glabrous, or sparsely cibate, shortly glandular-fimbriate. Bracts at or above middle of peduncle. Flowers c.1–5 cm, dark violet or white, fragrant. Sepals ovate, obtuse. Spur c.6 mm, exceeding calycine ap­pendages. Stigmatic beak vertical, its length equaling diameter of style. Capsule pubes­cent (Bobrov etal., 1934–1964; Rechinger, 1963–2001; Tutin etal., 1964; Davis, 1965– 1988; Sell and Murrell, 2006).
Vitex agnus-castus L., Sp. Pl.: 638 (1753)
Synonyms: Agnus-castus alba Carrière, Agnus- castus macrostachya Carrière, Agnus-castus robusta (Lebas) Carrière, Agnus-castus vulgaris Carrière, Vitex agnus Stokes, V. agnus-castus f. caerulea (Rehder) Moldenke, V. agnus-castus
subsp. haussknechtii Bornm., V. agnus-castus f. laciniosa (Ces.) Moldenke, V. haussknechtii (Bornm.) Prain, V. hyb ri da Moldenke, V. inte- gra Medik., V. latifolia Mill., V. lupinifolia Salisb., V. pseudonegundo (Hausskn.) Hand.- Mazz., V. robusta Lebas, V. verticillata Lam.
Family: Lamiaceae (Labiatae).
Arabic name: Fanjankosh.
Fig. 2.76. Viola odorata L. (By Fritz Geller-Grimm, CC BY-SA 2.5, https://upload.wikimedia.org/ wikipedia/commons/7/7a/Viola_odorata_fg01.JPG.)
Persian name: Panjangosht.
English name: Chaste tree, Vitex, Abra­ham’s balm, Lilac chastetree, Monk’s pepper (Fig. 2.77).
Distribution
Africa: Algeria, Egypt, Libya, Morocco, Tu­nisia. Asia: Afghanistan, Bahrain, Caucasus, Iran, Iraq, Lebanon, Oman, Pakistan, Syria, Tajikistan, Transcaucasus, Turkmenistan, Turkey, Yemen. Europe: Albania, Bulgaria, Cy­prus, France, Greece (Aegean Islands), Italy, Spain, Ukraine. Americas: Brazil, Cuba, Do­minican Republic, Honduras, Mexico, Peru, Puerto Rico, USA.
Botany, Collection, and Cultivation 111
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C.Y. Cheng & M.J. Liu, Z. chinensis Spreng.,
Z. flexuosa Wall., Z. jujuba f. allochroa C.Y.
Cheng & M.J. Liu, Z. jujuba f. apyrena C.Y.
Cheng & M.J. Liu, Z. jujuba f. carnosicalycis
C.Y. Cheng & M.J. Liu, Z. jujuba f. hetero-
formis C.Y. Cheng & M.J. Liu, Z. jujuba f.
tortuosa C.Y. Cheng & M.J. Liu, Z. melanog-
ona Bojer, Z. natsme Siebold, Z. nitida
Roxb., Z. officinarum Medik., Z. sativa
Gaertn., Z. sinensis Lam., Z. soporifera
(Lour.) Duhamel, Z. sororia Schult., Z. spi-
nosa (Bunge) Hu ex F.H. Chen, Z. trinervia
Roth, Z. vulgaris Lam., Z. vulgaris var. spi-
nosa Bunge, Z. zizyphus (L.) H. Karst.,
Zizyphon jujubum (Mill.) St.-Lag.
Family: Rhamnaceae.
Fig. 2.77. Vitex agnus-castus L. (By Krzsysztof Ziarnek, Kenraiz, CC BY-SA 4.0, https://upload. wikimedia.org/wikipedia/commons/9/94/Vitex_ agnus-castus_kz2.jpg.)
Morphology
Fragrant plant, 1–6 m tall. Young twigs with 4 obtuse angles, gray-puberulent. Leaves petiolate, with 5–7 stipitate leaflets; leaflets 1.5–10 cm, linear-lanceolate, acuminate, entire, whitish-tomentose be­neath, glabrous above. Calyx and outside of corolla tomentose. Corolla 8–10 mm, blue or pink. Drupe globose, reddish-black, slightly longer than calyx (Engler and Prantl, 1897–1915; Rechinger, 1963–2001; Davis, 1965–1988).
Ziziphus jujuba Miller, Gard. Dict. ed. 8:
no. 1 (1768)
Synonyms: Girtanneria jujuba (Mill.) Neck., Jububa mediterranea Bubani, Mansana ar­borea J.F. Gmel., Paliurus lucidus Carrière, Rhamnus circumcissa Russell ex Wall., R. lu­cida Salisb., R. mauritiana Soy.-Will., R. sop­orifera Lour., R. vulgaris Pers., R. zizyphus
L., Ziziphus acidojujuba C.Y. Cheng & M.J. Liu, Z. acidojujuba f. granulata C.Y. Cheng & M.J. Liu, Z. acidojujuba f. infecunda C.Y. Cheng & M.J. Liu, Z. acidojujuba f. trachysperma
Arabic name: Oʿnāb.
Persian name: Oʿnāb.
English name: Jujube, Jujuba, Chinese date
(Fig. 2.78).
Distribution
Africa: Algeria, Burkina Faso, Cameroon,
Tunisia, Morocco, Laos, Libya. Asia: Af-
ghanistan, China, Cyprus, India, Iran, Iraq,
Japan, Kazakhstan, Korea, Kyrgyzstan,
Lebanon, Maldives, Mongolia, Pakistan,
Syria, Tajikistan, Turkey, Turkmenistan, Uz-
bekistan. Europe: Bulgaria, France, Greece,
Italy, Romania, Spain. Americas: Cuba, Do-
minican Republic, Honduras, Jamaica, USA,
Venezuela.
Morphology
Small tree, rarely shrub, deciduous, to 10 m
tall, spinose or unarmed. Bark brown or gray-
brown, with long reduced branches, without
buds; branchlets (new branches) purple-red
or gray-brown, flexuose, smooth, with 2 stipu-
lar spines or not; long spines erect, to 3 cm,
stout; short spines recurved, developed from
old branches; annual branchlets pendulous,
green, solitary or 2–7-fascicled on short
shoots. Stipular spines slender, caducous;
petiole 1–6 mm, or to 1 cm on long shoots,
glabrous or sparsely puberulent; leaf blade
112 Farsad Nadjafi and Mohammad Reza Kanani
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Cultivation Methods of Some
Important Medicinal and Aromatic
Plants of Iran
Saffron (Crocus sativus)
Crocus sativus L. is a perennial stemless herb, commonly known as saffron. The dried stig­mas constitute the saffron of commerce. As cultivation of this species requires intensive labor and time, and the stigmas constitute a small part of the plant, it is the most expen­sive spice in the world. Cultivation of this expensive traditional spice is distributed from the Mediterranean region to Europe as well as to Asia. Iran is the world’s principal producer and exporter of saffron (85% of the worldwide saffron production). Due to very specific climate requirements, saffron grows in only a few countries. The main pro­ducer countries are Iran with 67,297 ha and a production amount of 239 t in 2010 and then India with 3200 ha cultivation area
CC BY-SA 4.0, https://commons.wikimedia.org/w/ index.php?curid=69284063.)
and a production amount of 12.5 t. The main producers in Europe are Greece with a culti­vation area of 1,750 ha and total production
of 7 t (in 2010) and Spain with 165 ha with a abaxially pale green, adaxially dark green, ovate, ovate-elliptic, or elliptic-oblong, 3–7 cm × 1.5–4 cm, papery, abaxially ± puberu­lent on major veins or glabrous, adaxially glabrous, 3-veined from base, base slightly asymmetric, subrounded, margin cren­ate-serrate, apex obtuse or rounded, rarely acute, mucronulate. Flowers yellow-green, bisexual, 5-merous, glabrous, solitary or 2–8 crowded in axillary cymes, shortly peduncu­late. Pedicel 2–3 mm. Sepals ovate-triangular, adaxially distinctly keeled. Petals obovate, about as long as stamens, clawed at base. Disk orbicular, thick, fleshy, 5-lobed. Ovary basally slightly immersed in disk; style 2-cleft to half. Drupe red at maturity, turn­ing red-purple, oblong or narrowly ovoid, 2–3.5 cm, (0.5–)1.5–2 cm in diameter; meso­carp fleshy, thick, sweet- or sour-tasting; stone acute or obtuse at both ends, 2­loculed, 1- or 2-seeded; fruiting pedicel 2–5 mm or longer. Seeds compressed-orbicular, c.1 cm × 0.8 cm (Rechinger, 1963–2001; Davis, 1965–1988).
total production of 3.2 t. Between countries,
Spain had the highest average yield per hec-
tare because of good agricultural practices
while the average yield in Iran was 3.5 kg/ha
(UNIDO, 2014). More than 96% of saffron
in Iran is being produced in Razavi Khorasan
and South Khorasan provinces. Saffron is
valued for its golden coloring, flavoring, and
aromatizing, but recently it is also being
considered by the pharmaceutical industry
for its antidepressant and anti-Alzheimer
activities (UNIDO, 2014).
Although about 150 volatile and non­volatile compounds have been detected in saffron stigmas, only less than 50 constitu ents have been identified so far (Winterhalter and Straubinger, 2000). Terpenes, terpene alcohols, and their esters represent the main components in volatile oil, among which sa­franal is the major component. Various cro­cins, crocetin, picrocrocin, and flavonoids like quercetin and kaempferol are the main non­volatile compounds (Liakopoulou-Kyriakides and Kyriakidis, 2002). The characteristic
-
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color of saffron is due to the presence of cro­cins, which are glucosyl esters of crocetin. The bitter taste of the spice is related to picrocrocin, a glycoside of safranal (Kanakis etal., 2004).
Saffron is an autumnal flowering geoph­ite characterized by a long summer rest in which the plant survives periods of drought by means of corms. Its biological cycle starts with its above-ground vegetative growth at the first autumn rains with the emission of leaves and flowers almost immediately and ends with the production of replacement corms in about 220 days. Unlike many other species of the Crocus genus that flower in the winter–spring period, in saffron flowering can occur from mid-October to the end of No­vember, essentially depending on the climatic conditions. It generally starts from 60 to 90 days after planting, mainly depending on sowing time, but snowy and cold periods may retard flowering. Saffron prefers direct sun exposure, even though in India it is cultivated together with almond trees. According to Fernández (2004) and Mollafilabi (2004), the best climatic conditions for high yields are rainfall in the autumn, warm summers, and mild winters. Saffron grows on a wide range of soils. Skrubis (1990) reported that the best yields are achieved on well-drained clay­calcareous and deep soil. Fernández (2004) suggests that clay is a good soil for saffron.
Saffron is cultivated in a wide range of environments with mild to dry climates. This plant is propagated by corm and the best cultivation time is in summer (end of July) when the plant is in dormancy period (Gresta et al., 2008a). Before planting, the soil should be completely cleared of weeds, ploughed at a depth of 25–30 cm, and left to rest from a few weeks to the entire winter. The application of about 20–30 t of organic manure per hectare is the most common fer
­tilization practice all over the world (Koockeki, 2004). The best corm size is 8–10 g and the best plant density is 19–20 t of corms per hectare (Koocheki etal., 2011). Koocheki et al. (2011) reported that best planting depth is 15 cm. They mentioned that the best planting method is row crop­ping with between- and inter-row distances of 15 and 20 cm, respectively (Koocheki
etal., 2009). Perennial crop techniques have highly variable durations from place to place: from 3–4 years in Spain, 6–8 years in India and Greece, and up to 12 years in Morocco (Ait-Oubahou and El-Otmani, 1999). With the aging of the saffron field, generally after 4–5 years, spice production declines because of increasing competition for water and nu­trients, fungal infection due to overcrowd­ing (Sampathu etal., 1984), and the reduced size and reproduction capability of corms. In a 10-year experiment, Grilli Caiola (2005) observed that corms left in the soil without management techniques continue produ
­cing daughter corms for up to 3–5 years but afterwards they degenerate and are no longer able to reproduce vegetatively. Water requirements of saffron are low and can be satisfied by the scarce rainfall when culti­vated in semiarid conditions. Researches in Iran (Koocheki, 2004) showed that this plant needs up to 3000 m
3
of irrigation per year. The most crucial moment for irrigation is after summer to awaken the corms but this coincides with autumn rains, so, except­ing a severe drought season, this may be considered unnecessary. Weeds are probably the main problem for saffron, since it is un­able to compete, above all because of its very low height. Nevertheless, due to the short time between the corm planting and flower­ing, severe weed problems start the follow­ing spring in perennial cultivation. From flowering, weeds can be left to grow until May and then cut without damaging the crop when the saffron leaves wither. Weeds are controlled mostly by hand. Gathering saffron flowers requires care and intensive manual labor: the flowers grow only a few centimeters above ground and, depending on vegetative activity, might be surrounded by several leaves that must not be damaged otherwise daughter corms will not be pro­duced. Flowers are harvested manually. About 350–450 labor-hours are needed to harvest 1 kg of the spice (Gresta et al., 2008b) (Fig. 2.79).
The saffron flower is highly ephemeral; given its very short life, it should be picked the same day of flowering and placed in bas­kets. The best practice is to pick the flower early in the morning each day, when the
114 Farsad Nadjafi and Mohammad Reza Kanani
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corolla is still closed, thereby preventing the stigmas from losing color and quality, avoid­ing any sudden deterioration by wind or rain (Zanzucchi, 1987; Tammaro, 1990) and al­lowing a ready separation into their con­stituent parts. After harvest, stigmas must be separated from the petals and stamen as soon as possible by opening the corolla and cutting the stigmas with the fingers below the branching where the style changes color (from red to yellow). Drying and storage methods are very important because a poor undertaking of this procedure can com­pletely compromise qualitative features of saffron (Carmona etal., 2005). According to the ISO norm, the moisture content may range between 10 and 12% w/w (ISO-3632­2:2003, 2003). The best drying temperature is between 35 and 40°C (Gresta et al ., 2008b). Generally, 1 ha of saffron may produce 10–15 kg of dried stigmas, but it can range widely, depending on the above mentioned factors, from 2 to 30 kg. In Iran, the average yield of saffron is around 5.4 kg/ha (Behzad
et al., 1992). In New Zealand, McGimpsey etal. (1997) reached yields of 24 kg/ha for
dried stigmas. The worst enemies of saffron are rodents and fungi (Tammaro, 1990; Go­liaris, 1999). Moles, rats, and rabbits can easily damage corms or eat leaves. Fungal at­tacks are mostly promoted by humid condi­tions. To avoid fungal infection, the best practices are crop rotation along with the re­moval and burning of infected plants.
Cumin (Cuminum cyminum L.)
Cumin (Cuminum cyminum L.) is an import­ant medicinal plant in Iran that has a long history of cultivation in the country. Cumin seeds are known for their spicy aroma and medicinal use (Hassanzadehdelouei et al.,
2013). This plant is cultivated in vast area in Iran, mostly in dryland agricultural systems. The area under cultivation and production of this plant in Iran have grown significantly in recent years. Today, the cultivation area is near 18,000 ha, with Razavi Khorasan Pro­vince (Fig. 2.80) accounting for more than 12,000 ha (Sefidkon, 2021).
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This species is a small bushy plant be­longing to the Apiaceae family with a height of 20–30 cm and a greenish color at early stage and a grayish color at maturity (Fig. 2.81).
The fruits are light brown or grayish in color and ovate or fusiform in shape (Hussein and Batra, 1998). The main producers of this plant are India, Iran, Syria, Egypt, and