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66 Theory and Practice of Squint and Orthoptics
approximation to their geometric locations so that a receptor 5° temporal to the fovea in one eye corresponds to a receptor 5° nasal to the fovea in the other eye.
Images falling on corresponding points give rise to a single mental impression to visual direction that we call sensory fusion. Corresponding retinal elements, therefore, have a common visual direction and permit a single visual impression. They do not permit double vision. The mental image resulting from the fusion of the two retinal images can be conceived as the image or mental image received by a cyclopean eye. This is an eye, presumably located in the middle of the forehead, that sees in visual space the fused images of the two individual eyes (Fig. 4.4).12 Normal retinal correspondence is thus the basis for normal binocular single vision. Thus the existence of corresponding retinal elements with their common relative subjective visual directions is the essence of binocular vision. It may be called the law of sensory correspondence in analogy with the law of motor correspondence.
If, during fixation of an object, one eye of the observer would be passively turned with forceps, the object would no longer be imaged on corresponding retinal points. While the images would fall on the fovea in the one eye, it
Fig. 4.4 Normal retinal correspondence. The visual direction
of f1 and f2 proceed from F the fovea of the imaginary cyclopean eye. A and B represent the points from which the visual direction belonging to a and a´, b and b´ would proceed (From Bielschowsky12).
would be received somewhere on the peripheral retina in the deviated eye. Since these two areas are non-corresponding and have different spatial values, diplopia would be present (i.e. the object would be seen double).
The oneness of the directional sensory
responses originating in each eye is impressively demonstrated by means of afterimages. If one
creates an afterimage on the retina of one eye, it will appear in the binocular field of view in the common visual direction of the stimulated retinal area and in its nonstimulated partner in the other eye. It is difficult, indeed almost impossible, for the observer to judge which eye carries the afterimages. It will continue to be seen and localized in the same direction, whether the eyes are open or closed or whether the stimulated eye is closed and the other eye held open.
If a horizontal afterimage is formed in one eye by a strong horizontal light stimulus, leaving the fovea unstimulated, and if a similar vertical afterimage is created in the other eye, the resulting visual percept is an afterimage in the form of a cross with a gap in its center (Fig. 4.5). The gap is seen because of the lack of stimulation in the foveae. The center of the horizontal and vertical afterimages is consequently a single spot localized in the principal common visual direction. The horizontal and vertical legs of the afterimages are oriented accordingly. It is of great importance to understand clearly that the appearance of the afterimage cross is independent of the position of the eyes. Once a lasting stimulus, such as an afterimage, has been imparted, its localization in subjective space depends solely on the visual direction of the retinal elements involved. One may topically anesthetize one eye and move it passively with a forceps or push it in any direction with one's finger-the cross remains a cross. No change in the relative localization of the vertical and horizontal afterimage will occur. The use of afterimages has an important place in the diagnosis of anomalous retinal correspondence. The principles underlying afterimage testing must be fully understood to guard against gross errors in interpretation.
Binocular Vision
A
Fig. 4.6 Diagrammatic representation of the theoretical
horopter (Vieth-Muller horopter circle). Points F, A and B fall on geometrically corresponding retinal points f1 and f2, a and a2 and b1 and b2, respectively.
67
1
B
Fig. 4.5 (A) Afterimages produced in the right and left eye,
respectively. The fovea is represented by the break in the afterimage. (B) The combined binocular afterimage forms a cross. The two gaps appear single.
HOROPTER
The term horopter, which literally means the horizon of vision, was introduced by Aguilonius.13 As we know, when, in normal binocular vision, both eyes fixate the same object point, this point will be imaged on corresponding retinal elements of the two eyes—the foveae. At the same time, there will be other object points in space besides the fixation point that also will be imaged on corresponding retinal elements of the two eyes. If the position of these other points is determined theoretically, a circle will be found passing through the fixation point and the entrance pupils of the two eyes. This circle is known as the theoretical or geometric horopter (Vieth-Muller horopter).
14,15
Theoretically, any point on it will stimulate corresponding retinal elements in the two eyes, because every point
on the horopter will form an angle with the two entrance pupils that is equal to the angle formed by the fixation point and the two entrance pupils (Fig. 4.6). It follows that there is a different horopter for each fixation distance.
In other words, the horopter can be defined as the sum total of points in physical space that stimulate corresponding retinal elements of the two eyes. It is a complex mathematical model having all three dimensions. The longitudinal horopter is that surface that determines horizontally distributed object points in space. The longitudinal horopter curve is a line formed by the intersection of the visual plane and the longitudinal horopter plane. The visual plane is determined by the fixation point and the centres of the two pupils.
16, 17
The horopter can be determined with an instrument utilizing movable, vertically placed wires set at a certain distance. While fixation is maintained upon a central wire, those on either side in the periphery are adjusted forward or backward by the subject until they appear to lie in the same visual plane. In determining the horopter, the difference between subjective visual space and physical space is again
13
68 Theory and Practice of Squint and Orthoptics
demonstrated. Subjectively, the movable wires appear to lie in the same visual plane; however, they actually lie on a curved surface. Thus, the horopters found experimentally do not actually coincide with the theoretical horopters. They are called empirical horopters.
PHYSIOLOGIC DIPLOPIA
Since only object points which lie on the horopter will be imaged on corresponding retinal elements, all objects that are either nearer or farther away than the horopter will be imaged on disparate (i.e. non-corresponding) retinal elements and, consequently, will be seen double.
Point a in Fig. 4.7A, located closer than the horopter (point b), is imaged temporal to the fovea in each eye. The right eye will see this point as being on the left side of the fixation
point, while the left eye will see it to the right of the fixation point. In other words, point a will be seen double (a1 and a2). Since each eye perceives it as being on the opposite side, this is referred to as crossed diplopia. With respect to the retina, point a shows temporal disparity. Conversely point a, which is located beyond the horopter (point b) in Fig. 4.7B; will be imaged on the nasal retinae of both eyes and, therefore, will also appear double. However, this time the diplopia will be uncrossed—the right eye will localize the object to the right of the fixation point, while the left eye will place it to the left. The retinal images will have nasal disparity.
Physiologic diplopia can be demonstrated by holding two pencils vertically in front of the eyes with one pencil about twice as far away as the other. If either one of them is fixated, the other will be seen double. If the more distant one is fixated, crossed diplopia will be experienced. If the closer one is fixated, the diplopia will be uncrossed.
This demonstration will not be immediately successful in all subjects, since one of the images may be suppressed from conscious perception, a process which also is physiologic, as is explained later in this chapter.
In view of what has been said about the horopter and physiologic diplopia, it should be expected that one would have constant diplopia during casual daily vision, with the exception of the object of fixation and the few objects that happen to be on the horopter for the momentary fixation distance. Obviously, this is not the case; and it is not primarily suppression that prevents the occurrence of physiologic diplopia, but the exclusiveness with which our attention is usually directed to the fixated object. Although diplopia is present, this rarely enters consciousness. Another contributing factor is the low visual acuity of the peripheral retina, which frequently does not yield sufficient resolution of peripherally located objects for the recognition of diplopia.
Fig. 4.7 Diagram illustrating physiological diplopia. Points
a and b which are not located on the horopter, are imaged on non-corresponding retinal points and so physiological diplopia either crossed (A) or uncrossed (B) is produced.
BINOCULAR FUSION
When images of an object fall on corresponding retinal points, in the normal subject, they seem to be fused into a single mental impression.
Binocular Vision
69
Sensory fusion, i.e. single vision is the hallmark of retinal correspondence. Put otherwise, the
stimulus to sensory fusion is the excitation of corresponding retinal elements. It should be distinguished from motor fusion, which refers to the ability to align the eyes in such a manner that sensory fusion can be maintained. The stimulus for these fusional eye movements is retinal disparity. Unlike sensory fusion, the motor fusion is exclusive function of the extra­foveal retinal periphery. No stimulus for motor fusion exists when the images of a fixated visual object fall on the fovea of each eye.
Under normal conditions, sensory fusion occurs when corresponding retinal elements of the two eyes are stimulated by images from the same object. Thus for sensory fusion to occur, the images not only must be located on corresponding retinal areas but also must be sufficiently similar in size, brightness, and sharpness. Unequal images are a severe sensory obstacle to fusion. Obstacles to fusion may become important factors in the etiology of strabismus. However, fusion will also take place when the stimuli in the two eyes are not identical but only similar. The dissimilarity may be one of form, size, colour, luminosity, distinctness or contrast. To a limited extent, it is also possible to fuse signals from non-corresponding points (see discussion on fixation disparity below).
SENSORY FUSION
Sensory fusion, also known as binocular fusion, thus, can be defined as the ability of the brain to combine the slightly disparate images from each eye into a single, unified visual perception. This process is crucial for achieving and maintaining binocular vision, which allows for depth perception and a three-dimensional perception of the world.
The anatomic basis which allows sensory fusion to occur is the course of the visual nerve fibres.
By crossing in the chiasma, nerve fibres from the nasal retina are brought to the same side as their counterparts from corresponding points of the temporal retina of the other eye. While ascending in the visual pathway, the fibres from corresponding retinal points converge until finally they are adjacent to each other. They terminate in the same cortical cell complexes.
But for our knowledge of the anatomic arrangement, little definite information exists on the physiology of sensory fusion. It is probable that fusion occurs in the cortex. It is not certain, however, whether the two signals transmitted separately from two corresponding points are synthesized so that they simultaneously contribute to the resulting single perception, or whether the two signals are mutually exclusive and are utilized alternately to be combine into the resulting single constant perception so that first one eye and then the other contributes its stimulus to a given point of the whole.
MOTOR FUSION
As mentioned above, the motor fusion is a
corrective vergence movement in response to
image disparity.
Motor fusion adjusts eye position to maintain
sensory fusion.
As a fixation target approaches the observer,
the retinal images move temporally from each
fovea if the eye remains in an unchanged
position.
To prevent diplopia, the crossed image
disparity induces both eyes to converge (turn
inwards) and maintain the image focused on
the foveae.
A similar divergent movement occurs as
objects move from near to far.
Fusional reserve indicates the level at which
motor fusion breaks down, usually causing
diplopia. It can be measured by adding prism
bars (base in or out) until fusion is lost
(Table 4.1).
Torsional fusional vergences also exist for up
to 6-10° of torsional image disparity.
At birth, coordinated conjugate eye movements are absent, visual perception is poor, and fusion is not established as a stable binocular function.
If development proceeds normally, fusion evolves later as a conditioned reflex. If there is a significant abnormality in the optical performance of the eyes, in the function of the nerve elements that are concerned with the transmission or perception of visual stimuli, or in the motor cooperation of the two eyes, fusion
70 Theory and Practice of Squint and Orthoptics
Table 4.1 Normal fusional amplitudes
Testing distance (m) Convergence Divergence Vertical
(prism diopters, ) (prism diopters, ) (prism diopters, )
6 (distance) 16 6 5–6
0.25 (near) 32 16 3–4
may never develop normally. It is conceivable that there may be individuals in whom a basic ability for fusion does not exist.
PANUM’S AREA
As stated previously that only object points which lie on the horopter and which stimulate corresponding retinal points in the two eyes are seen as single is not quite exact and must be qualified. When a horopter is determined experimentally, it is found that an object may be located a certain distance in front of or behind it without producing diplopia. The field in front of and behind the horopter, in which the expected diplopia does not occur is known as Panum’s fusional space.
Panum’s fusional space is smallest at the fixation point (Fig. 4.8). Toward the periphery, it gains increasingly in depth, so that objects located peripherally may be farther from the horopter without producing diplopia than objects located more centrally. Any point not on
the horopter is not imaged on corresponding retinal areas but will stimulate disparate retinal areas. If, despite the disparity, no diplopia occurs, fusion of signals from non-correspon­ding points must have taken place. Thus, a signal from a given retinal point not only is fused with the signal from its corresponding point in the other eye, but also may be fused with signals from a limited area surrounding that corresponding point. Such a retinal Panum’s area corresponds in size to the depth of Panum’s fusional space. Since the latter is small in the vicinity of the fixation point, Panum’s areas in the fovea are also small, actually in the order of 5’ of arc. Away from the fixation point, Panum’s fusional space enlarges (i.e. the size of Panum’s areas in the peripheral retina increases). Since retinal disparity in the horizontal meridian can be overcome to a larger extent than disparity in the vertical meridian, Panum’s areas have the shape of an oval with the longer axis horizontal. If the fixation distance is more than 20 m, objects lying behind the horopter will always be seen as single, since the disparity of their images is always smaller than Panum’s areas.
Fig. 4.8 Diagram of Panum's fusional space.
FIXATION DISPARITY
Under laboratory conditions, it can be demonstrated that, during binocular fixation, the point of fixation is rarely ever imaged exactly on corresponding points of the two foveae but that the primary line of sight of one eye misses the fixation point very slightly, being either under-converged or over-converged (Fig. 4.9). This phenomenon is called fixation disparity. It does not give rise to diplopia, because the disparity with which the fixation point is imaged on the two retinae is less than the size of Panum’s area.
Thus, in fixation disparity, one eye will fixate the object directly with the central fovea, while the other eye will fixate slightly eccentrically.
Fig. 4.9 Diagram illustrating fixation disparity.
However, the horopter simply passes through the actual point of fixation rather than through the fixation stimulus. Fixation disparity may arise from a small foveal scotoma or from an oculomotor imbalance which is tending to pull the eyes away from the appropriate angle of convergence. An example of a horopter showing fixation disparity was obtained by Olge18 on his own eyes.
DICHOPTIC STIMULATION
Dichoptic stimulation simply refers to a different stimulation in the two eyes, which occurs when binocular stimuli fall on non-corresponding points on the two retinae. There are five classes of percepts which are obtained, depending upon the degree of non-correspondence between the stimuli:
1. Depth with fusion;
2. Depth with diplopia;
3. Diplopia without depth;
4. Binocular rivalry and suppression; and
5. Binocular lustre.
DEPTH WITH FUSION AND DEPTH WITH DIPLOPIA
Strictly speaking, these are not generally referred to as dichoptic stimuli, since the two retinal patterns are sufficiently similar to be
Binocular Vision
71
combined into a unified impression (particularly for fused stereopsis). Therefore, these will be dealt in the section of stereopsis.
DIPLOPIA WITHOUT DEPTH
When the degree of non-correspondence is such that no fusion occurs and diplopia results without depth perception. Depending upon the location of non-corresponding points stimulated, the diplopia may be homonymous or uncrossed (Fig. 4.7B) and heteronymous or crossed (Fig. 4.7A). Sensory adaptations which may occur to compensate for the annoying diplopia include: suppression, amblyopia, and abnormal retinal correspondence. Motor mechanisms which compensate for diplopia are an abnormal head posture and certain permanent changes in the extraocular muscles such as inhibitional palsy, overaction and contractures.
RETINAL RIVALRY AND SUPPRESSION
When dissimilar contours are presented to corresponding retinal areas (having same visual direction), fusion becomes impossible, since two dissimilar objects localized in the same place give rise to a conflict and confusion. The conflict which results when contradictory and incompatible signals are transmitted from corresponding points to the visual cortex is termed retinal rivalry. The phenomenon of retinal rivalry, also termed as binocular rivalry, must be clearly distinguished from local adapta­tion, or Troxer’s phenomenon.24 Binocular rivalry may also be produced by uniform surfaces of different colour (colour rivalry) and unequal luminances of the two targets. Many combinations of contours, colours and luminances have been studied exhaustively since the days of Panum,25 Fechner, Helmholtz27 and Hering.


Retinal rivalry is a very normal event, since
28
corresponding areas are consistently exposed to dissimilar images from objects located outside Panum’s fusional space.
The visual system responds to retinal rivalry by rejecting the signals transmitted from a given area of one retina in favour of those coming from
26
72 Theory and Practice of Squint and Orthoptics
the corresponding parts of the other retina. Thus, only information from one retina becomes a visual perception while the information simultaneously transmitted from corresponding parts of the other retina is not used, so that the involved area may be considered temporarily blind. This reaction to the influx of conflicting information is called suppression.


Suppression is an innate, involuntary process. Under certain conditions, all the signals from one retina may be suppressed so that no information from that eye reaches the threshold of conscious perception.
29-33
Usually, however, responding to the demand of the momentary conditions, suppression is restricted to limited areas in both retinae so that those portions of the subjective visual field in which fusion cannot occur may be composed, like a mosaic, of pieces of information from only one or the other retina in constant variation (Fig. 4.10).
Although an eye may be suppressed for long periods, suppression basically is a transient phenomenon, present only momentarily when needed. Because it is a physiologic function that does not have to be learned but is readily available, it frequently becomes the firstline of defence against pathologic interruption or
Fig. 4.10 Retinal rivalry produced by dissimilar contours:
(A) pattern viewed by left eye; (B) Pattern viewed by right eye; (C) binocular impression (mosaic of pieces). After Panum.25
embarrassment of bifoveal single vision. For example, a patient who has a marked refractive error in only one eye may show suppression of that eye to eliminate the disturbing effect of the blurred image. In patients with strabismus, a condition in which objects are not imaged on normally corresponding retinal areas, suppre­ssion is a frequent response to solve the problem of incompatible information transmitted from corresponding areas of the two eyes. It has been shown experimentally that retinal rivalry and the suppression resulting from it are influenced by certain conditions. If non-fusable patterns are presented to the two eyes, it is found that perception from corresponding areas alternates equally between the two eyes, if the objects are of equal brightness, size and prominence. The rate of alternation under such test conditions is influenced by the brightness of the field, the size of the target area, the distinctness of the targets, and the location of the stimulated area in the retina. If any of these factors is impaired equally for both eyes, the rate of alternation will decrease. If the visual conditions are improved, it will increase. However, suppression will always alternate equally between the two eyes.
In contrast, by changing the stimulus conditions for only one eye, the rate of alternation can be changed markedly in its favour or disfavour.
Retinal or binocular rivalry fluctuations are similar in many respects to fluctuations of attention, and are widely supposed to be under voluntary control to some extent. Actually, a number of studies have found that there is very little voluntary control over which eye dominates at any given time.34 The change of dominance is not affected by eye blinks35 or by variations in accommodation or pupil size.36 In fact, the fluctuations in rivalry are well described by a sequentially independent random variable with no periodicities, as though the arrival of each change in dominance had no effect on the occurrence of subsequent changes.
37
The phenomenon of retinal rivalry has been explained in neurophysiologic terms by the presumption that separate channels are present for the right and left eyes which compete for access to the visual cortex. A third binocular
Binocular Vision
73
channel is activated only by fusible input.
38,39
Because of this competition and inhibition elicited, only fragments of the image seen by each eye are transmitted to the visual cortex in the case of non-fusible binocular output. This hypothesis looks logical but needs experimental verification.
BINOCULAR LUSTRE
The lustrous appearance of surfaces like a waxed surface, tabletop or a car body is essentially due to binocular lustre. It results from the different position of partially reflected objects in the surface by virtue of the different position of the two eyes. The partial reflection provides a fixation place at which the partially reflected image usually has a large disparity and hence areas of binocular luminance difference. The lustrous region is not localizable in depth, but it seems unitary and does not fluctuate in the manner of binocular rivalry. Binocular lustre was described by early authors in visual science, such as Panum19 and Helmholtz,21 as a kind of lustrous or shimmering surface of indeterminate depth.
The detection of binocular lustre during static and dynamic random dot stereogram testing is even more rapid than the detection of depth changes.
40-42
STEREOPSIS
Stereopsis is the visual appreciation of three dimensions during binocular vision. It occurs through fusion of signals from disparate retinal elements.
As shown in Fig. 4.11, if one views two posts, A and B, situated at slightly different distances, with both eyes open, their separation in depth is readily apparent to a normal person. This perception of depth is maintained as long as both eyes are used. If either eye is closed, the effect of depth then vanishes. If the right eye is closed, the appearance of the two posts is that shown on the left hand side of Fig. 4.11, and, if the left eye is closed, the appearance is that shown on the right side of the figure. The only difference between these two images is that the posts seem to be farther apart in the frontoparallel plane when seen with the left eye
Fig. 4.11 Disparateness of retinal images producing
stereopsis.
than when seen with the right. This difference in lateral separation is due to the fact that the receptors stimulated in the left eye are more widely separated from each other than those in the right retina (note distance between a and b of two eyes in Fig. 4.11).
An object point which is located in front of the fixation point but is still within the limits of Panum’s fusional space will stimulate disparate retinal elements. The disparity will be temporal, each image being temporal to the point that corresponds to the location of the image in the other eye. Although both images are fused since they are within the limits of Panum’s area, the fact that they are temporarily desperate is not lost in the resulting single perception but gives the perception of its additional quality of nearness relative to the fixation point. Conversely, when an object is located behind the horopter but still within the Panum’s fusional space, nasal disparity will exist, adding the quality of farness to the perception. Thus, it can be seen from Fig. 4.12 that, when the images do not fall on exactly corresponding retinal
74 Theory and Practice of Squint and Orthoptics
Fig. 4.12 Scheme showing the effect produced by
stimulation of slightly incongruous retinal points. a-a', corresponding points; b-b' corresponding points; F-F', the foveas.
points, an impression that the image is in front or behind a given frontal plane is produced. In the figure 4.12, F and F' are the foveae fixating a point C, straight ahead. This point determines the frontal plane, XX’. An object A, whose image falls on a in the left eye produces an image at b’ in the right eye. b’ is not the corresponding point, which is a’. Because of this, the impression is that the point A lies nearer the eyes than the frontal plane, XX’. Similarly, an object O, whose image falls on b in the left eye produces its image at a' in the right eye. The corresponding retinal point of b is b’. The image of object O lies to the nasal side of this point at a’ in the right eye. The impression is gained, therefore, that point O lies behind frontal plane XX’.
Thus, it proves that in general nasal disparity gives the impression of remoteness of an object, and temporal disparity gives the impression of nearness of an object.
Stereopsis is a unique cognition, a distinct perceptional quality of its own. If one does not have it, one cannot learn it even in the presence of all requirements such as bifoveal fixation, fusion, and good visual acuity. Although stereopsis is the most important factor in judging distances, especially at close range, additional information about the absolute and
relative distances of objects is gained in several other ways. While in bifoveal vision these other means are only an addition to stereopsis, they are the only source of information on which a monocular observer can base his distance judgement.
Worth classified fusion in three degrees. First-
degree fusion was the simultaneous macular perception of two different non-fusable objects. Second-degree fusion was characterized as the ability of the motor system to align the eyes in such a way that sensory fusion was possible. Third-degree fusion was stereopsis. Worth’s classification of third-degree fusion, however, is not entirely correct since stereopsis is a separate quality of the visual system, a sensory responsiveness to disparate stimuli that can occur, under certain conditions, without second­degree fusion being present.
In a two-dimensional representation, certain
colours seem to stick out and others do not. Colour stereopsis is related to the differential refraction for various wavelengths of light. The different blur circles can stimulate disparate retinal points within Panum’s area and thus give true stereopsis to a flat picture in which there is no lateral or horizontal separation of the retinal images.
PHYSIOLOGICAL BASIS OF STEREOPSIS
Stereopsis: A function of spatial disparity
Stereopsis arises when horizontally disparate retinal elements are stimulated simultaneously. The fusion of such disparate images results in a single visual impression perceived in depth, provided the fused image lies within Panum's area of single binocular vision. Vertical disparities produce no stereoscopic effect. A solid object placed in the median plane of the head produces unequal images in the two eyes due to horizontal separation of the two eyes (Fig. 4.13). The sensory fusion of the two unequal retinal images results in a three­dimensional percept.
A stereoscopic effect can also be produced by two-dimensional pictures, some elements of which are imaged on corresponding retinal points to give the frame of reference for the relative localization of other elements of figures
43
Binocular Vision
75
Fig. 4.13 A solid object placed in the midline of the head creates slightly different or disparate retinal images, the fusion
of which results in a three-dimensional sensation.
constructed to provide horizontally disparate imagery. Such figures must be viewed separately but binocularly in a stereoscope or some haploscopic device.
recognition of a three-dimensional form. Julesz refers to this additional mechanism as global stereopsis, since the problem of ambiguities can only be resolved on a global basis. Therefore, a satisfactory theory for binocular depth
Local and global stereopsis
44,45
Julesz
studied stereopsis perception utilizing
perception must obviously take account of both local and global stereopsis.
random stereogram and put forward the concept of local and global stereopsis. The term local stereopsis is used to denote stereopsis elicited by the dot-by-dot or square-by-square matching process that occurs between the right and left stereogram.
When there is ambiguity as to which elements in the two retinal images correspond to each other, a global process is needed that evaluates different possible sets of corresponding pairs and selects one set of matched pairs that, by their depth values, can provide the data for
Fine versus coarse stereopsis
Fine stereopsis is a highly specific pattern matching process involving very local features of the two retinal images and operating over a relatively narrow range of spatial disparities probably no more than about 0.5 degrees.
Dissimilar images cannot be simultaneously perceived. If binocular vision is to occur, the two retinal images must be closely similar, otherwise there is retinal rivalry and suppression of one or other of the antagonistic image features.