Адаптация животных к хроническим факторам
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ɝɨɪɢɡɨɧɬɚɥɶɧɨɦɭ ɞɢɚɦɟɬɪɭ ɰɟɧɬɪɚɥɶɧɨɣ ɡɨɧɵ ɩɪɚɜɨɦɭ ɪɚɡɦɟɪɭ ɫɜɨɛɨɞɧɨɣ ɡɨɧɵ ɜɟɪɯɧɟɦɭ ɢ ɩɪɚɜɨɦɭ ɪɚɡɦɟɪɭ ɥɨɜɱɟɣ ɡɨɧɵ ɢ ɞɥɢɧɚɦɢ ɪɚɞɢɭɫɨɜ Ɇɟɠɞɭ -ɣ ɢ -ɣ ɜɨɡɪɚɫɬɧɵɦɢ ɫɬɚɞɢɹɦɢ ɩɚɭɤɨɜ ɪɚɡɥɢɱɢɹ ɞɨɫɬɨɜɟɪɧɵ ɩɨ ɬɟɦ ɠɟ ɩɨɤɚɡɚɬɟɥɹɦ ɢ ɩɨ ɜɟɪɯɧɟɦɭ ɪɚɡɦɟɪɭ ɫɜɨɛɨɞɧɨɣ ɡɨɧɵ ɜɫɟɦ ɪɚɡɦɟɪɚɦ ɥɨɜɱɟɣ ɡɨɧɵ ɩɨ ɱɢɫɥɭ ɥɨɜɱɢɯ ɧɢɬɟɣ ɫɩɪɚɜɚ ɢ ɫɧɢɡɭ ɋɟɬɢ ɩɚɭɤɨɜ ɧɚ -ɣ ɢ -ɣ ɫɬɚɞɢɹɯ ɞɨɫɬɨɜɟɪɧɨ ɪɚɡɥɢɱɚɸɬɫɹ ɩɨ ɫɥɟɞɭɸɳɢɦ ɩɨɤɚɡɚɬɟɥɹɦ ɜɟɪɯɧɢɣ ɩɪɚɜɵɣ ɢ ɧɢɠɧɢɣ ɪɚɡɦɟɪɵ ɥɨɜɱɟɣ ɡɨɧɵ ɢ ɜɫɟ ɞɥɢɧɵ ɪɚɞɢɭɫɨɜ Ɇɟɠɞɭ -ɣ ɢ -ɣ ɫɬɚɞɢɹɦɢ ɪɚɡɜɢɬɢɹ ɩɚɭɤɨɜ ɪɚɡɥɢɱɢɹ ɞɨɫɬɨ ɜɟɪɧɵ ɩɨ ɚɧɚɥɨɝɢɱɧɵɦ ɩɨɤɚɡɚɬɟɥɹɦ ɩɨ ɧɢɠɧɟɦɭ ɪɚɡɦɟɪɭ ɫɜɨɛɨɞɧɨɣ ɡɨɧɵ ɥɟɜɨɦɭ ɪɚɡɦɟɪɭ ɥɨɜɱɟɣ ɡɨɧɵ ɢ ɩɨ ɱɢɫɥɭ ɥɨɜɱɢɯ ɧɢɬɟɣ ɫ ɤɚɠɞɨɣ ɫɬɨɪɨɧɵ Ɇɟɠɞɭ ɫɟɬɹɦɢ ɩɚɭɤɨɜ ɧɚ -ɣ ɢ -ɣ ɜɨɡɪɚɫɬɧɵɯ ɫɬɚɞɢɹɯ ɜɵɹɜɥɹɸɬɫɹ ɞɨɫɬɨɜɟɪɧɵɟ ɪɚɡɥɢɱɢɹ ɩɨ ɬɚ ɤɢɦ ɩɨɤɚɡɚɬɟɥɹɦ ɤɚɤ ɜɟɪɬɢɤɚɥɶɧɵɣ ɢ ɝɨɪɢɡɨɧɬɚɥɶɧɵɣ ɞɢɚɦɟɬɪɵ ɰɟɧɬɪɚɥɶɧɨɣ ɡɨ ɧɵ ɜɫɟ ɪɚɡɦɟɪɵ ɥɨɜɱɟɣ ɡɨɧɵ ɢ ɜɫɟ ɞɥɢɧɵ ɪɚɞɢɭɫɨɜ ɚ ɬɚɤɠɟ ɱɢɫɥɨ ɥɨɜɱɢɯ ɧɢɬɟɣ ɜ ɤɚɠɞɨɦ ɫɟɤɬɨɪɟ Ɍɚɤɢɦ ɨɛɪɚɡɨɦ ɦɨɠɧɨ ɫɱɢɬɚɬɶ ɱɬɨ ɦɢɧɢɦɚɥɶɧɵɟ ɢɡɦɟɧɟɧɢɹ ɜ ɫɬɪɭɤɬɭɪɟ ɫɟɬɟɣ A. marmoreus ɩɪɨɢɫɯɨɞɹɬ ɩɪɢ ɩɟɪɟɯɨɞɟ ɨɬ -ɣ ɤ -ɣ ɥɢɱɢɧɨɱɧɨɣ ɫɬɚɞɢɢ ɪɚɡɜɢɬɢɹ ɚ ɧɚɢɛɨɥɶɲɢɟ ɩɪɢ ɩɟɪɟɯɨɞɟ ɨɬ ɩɨɫɥɟɞɧɟɣ ɥɢɱɢɧɨɱɧɨɣ ɫɬɚɞɢɢ ɪɚɡɜɢɬɢɹ ɤ ɩɨɥɨɜɨɡɪɟɥɨɣ ɨɫɨɛɢ
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ɋɪɟɞɧɢɟ ɡɧɚɱɟɧɢɹ ɩɨɤɚɡɚɬɟɥɟɣ ɥɨɜɱɢɯ ɫɟɬɟɣ |
Ɍɚɛɥɢɰɚ |
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ɩɚɭɤɨɜ A. marmoreus ɜ ɨɧɬɨɝɟɧɟɡɟ |
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ɉɨɤɚɡɚɬɟɥɢ |
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ɋɬɚɞɢɢ ɪɚɡɜɢɬɢɹ |
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3-ɹ |
4-ɹ |
5-ɹ |
ɩɨɥɨɜɨɡɪɟ |
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ɥɚɹ |
ɑɢɫɥɨ ɜɢɬɤɨɜ |
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ɰɟɧɬɪ Ɂɨɧɵ |
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ȼɟɪɬ ɞɢɚɦ ɰ ɡ |
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Ƚɨɪɢɡ ɞɢɚɦ ɰ ɡ |
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ȼɟɪ ɪɚɡɦɟɪ ɫɜ ɡ |
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ɉɪɚɜɵɣ |
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8 |
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ɪɚɡ ɫɜɨɛɨɞ ɡ |
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ɇɢɠɧɢɣ ɪɚɡɦɟɪ |
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ɫɜɨɛɨɞɧɨɣ ɡɨɧɵ |
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Ʌɟɜɵɣ ɪɚɡɦɟɪ |
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ɫɜɨɛɨɞɧ ɡɨɧɵ |
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ȼɟɪɯ ɪɚɡɦɟɪ |
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ɥɨɜɱɟɣ ɡɨɧɵ |
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ɉɪɚɜɵɣ ɪɚɡɦɟɪ |
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ɥɨɜɱ ɡɨɧɵ |
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ɇɢɠɧɢɣ ɪɚɡɦɟɪ |
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ɥɨɜɱɟɣ ɡɨɧɵ |
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30,4* |
Ʌɟɜɵɣ ɪɚɡɦɟɪ |
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ɥɨɜɱ ɡɨɧɵ |
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ȼɟɪɯ ɞɥɢɧɚ ɪɚ |
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4* |
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ɞɢɭɫɨɜ |
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ɉɪɚɜɚɹ ɞɥɢɧɚ |
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ɪɚɞɢɭɫɨɜ |
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24,8* |
ɇɢɠɧ ɞɥɢɧɚ |
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ɪɚɞɢɭɫɨɜ |
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26,1* |
Ʌɟɜɚɹ ɞɥɢɧɚ ɪɚ |
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ɞɢɭɫɨɜ |
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ɑɢɫɥɨ ɥɨɜɱɢɯ |
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1 |
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* |
ϲϲ
ɧɢɬɟɣ ɜɟɪɯ |
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ɑɢɫɥɨ ɥɨɜɱ ɧɢ |
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ɬɟɣ ɜɩɪɚɜɨ |
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ɑɢɫɥɨ |
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* |
ɥɨɜɱ ɧɢɬɟɣ ɜɧɢɡ |
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ɑɢɫɥɨ ɥɨɜɱ ɧɢ |
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* |
ɬɟɣ ɜɥɟɜɨ |
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ɑɢɫɥɨ ɪɚɞɢɭɫɨɜ |
6 |
6 |
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ɉɪɢɦɟɱɚɧɢɟ ȼɫɟ ɪɚɡɦɟɪɵ ɩɪɢɜɟɞɟɧɵ ɜ ɦɢɥɥɢɦɟɬɪɚɯ (*)-ɫɬɚɬɢɫɬɢɱɟɫɤɢ ɞɨɫ ɬɨɜɟɪɧɵɟ ɢɡɦɟɧɟɧɢɹ ɩɨɤɚɡɚɬɟɥɟɣ ɩɨ ɨɬɧɨɲɟɧɢɸ ɤ ɩɪɟɞɵɞɭɳɟɣ ɜɨɡɪɚɫɬɧɨɣ ɫɬɚ ɞɢɢ
ɋɪɟɞɢ ɜɫɟɯ ɢɫɫɥɟɞɭɟɦɵɯ ɩɨɤɚɡɚɬɟɥɟɣ ɥɨɜɱɢɯ ɫɟɬɟɣ ɩɚɭɤɨɜ ɜ ɩɪɨɰɟɫɫɟ ɨɧɬɨ ɝɟɧɟɡɚ ɬɨɥɶɤɨ ɞɜɚ ɢɡɦɟɧɹɸɬɫɹ ɜ ɫɬɨɪɨɧɭ ɭɦɟɧɶɲɟɧɢɹ ɱɢɫɥɨ ɜɢɬɤɨɜ ɜ ɰɟɧɬɪɚɥɶɧɨɣ ɡɨɧɟ ɢ ɱɢɫɥɨ ɪɚɞɢɭɫɨɜ ɉɪɢɱɟɦ ɩɨ ɜɬɨɪɨɦɭ ɩɨɤɚɡɚɬɟɥɸ ɞɨɫɬɨɜɟɪɧɵɯ ɪɚɡɥɢɱɢɣ ɧɟ ɜɵɹɜɥɟɧɨ ɋɥɟɞɨɜɚɬɟɥɶɧɨ ɱɢɫɥɨ ɪɚɞɢɭɫɨɜ ɹɜɥɹɟɬɫɹ ɜɚɪɢɚɛɟɥɶɧɵɦ ɩɨɤɚɡɚɬɟɥɟɦ ɫɟɬɢ ɧɨ ɧɟ ɞɨɫɬɨɜɟɪɧɨ ɢɡɦɟɧɹɟɬɫɹ ɜ ɢɧɞɢɜɢɞɭɚɥɶɧɨɦ ɪɚɡɜɢɬɢɢ ɩɚɭɤɨɜ ɇɚɢɛɨɥɟɟ ɩɨɫɥɟɞɨɜɚɬɟɥɶɧɨ ɢɡɦɟɧɟɧɢɹ ɷɥɟɦɟɧɬɨɜ ɩɨɜɟɞɟɧɢɹ ɨɬɜɟɬɫɬɜɟɧɧɵɯ ɡɚ ɪɚɡɦɟɪɵ ɰɟɧ ɬɪɚɥɶɧɨɣ ɡɨɧɵ ɥɨɜɱɟɣ ɡɨɧɵ ɢ ɞɥɢɧ ɪɚɞɢɭɫɨɜ Ɂɚɦɟɬɧɨɟ ɭɜɟɥɢɱɟɧɢɟ ɱɢɫɥɚ ɥɨɜɱɢɯ ɧɢɬɟɣ ɩɪɨɢɫɯɨɞɢɬ ɜɨ ɜɡɪɨɫɥɨɣ ɫɬɚɞɢɢ ɇɚɢɛɨɥɟɟ ɜɚɪɢɚɛɢɥɶɧɵɦɢ ɹɜɥɹɸɬɫɹ ɩɨɤɚ ɡɚɬɟɥɢ ɫɜɨɛɨɞɧɨɣ ɡɨɧɵ ȼɨ ɜɡɪɨɫɥɨɣ ɫɬɚɞɢɢ ɷɬɨɬ ɩɨɤɚɡɚɬɟɥɶ ɢɡɦɟɧɹɥɫɹ ɨɬ ɞɨ ɦɦ ɜɫɬɪɟɱɚɥɢɫɶ ɫɟɬɢ ɜ ɤɨɬɨɪɵɯ ɫɜɨɛɨɞɧɚɹ ɡɨɧɚ ɨɬɫɭɬɫɬɜɨɜɚɥɚ
Ɍɚɤɢɦ ɨɛɪɚɡɨɦ ɧɚ ɨɫɧɨɜɚɧɢɢ ɩɪɨɜɟɞɟɧɧɵɯ ɢɫɫɥɟɞɨɜɚɧɢɣ ɜɨɡɪɚɫɬɧɨɣ ɫɬɪɭɤ ɬɭɪɵ ɥɨɜɱɢɯ ɫɟɬɟɣ ɩɚɭɤɨɜ-ɤɪɭɝɨɩɪɹɞɨɜ A. marmoreus ɦɨɠɧɨ ɫɱɢɬɚɬɶ ɱɬɨ ɨɫɧɨɜ ɧɨɣ ɚɥɝɨɪɢɬɦ ɩɨɜɟɞɟɧɢɹ ɩɨɫɬɪɨɟɧɢɹ ɥɨɜɱɢɯ ɫɟɬɟɣ ɮɨɪɦɢɪɭɟɬɫɹ ɧɚ ɬɪɟɬɶɟɣ ɥɢɱɢ ɧɨɱɧɨɣ ɫɬɚɞɢɢ ɢ ɧɟ ɩɪɟɬɟɪɩɟɜɚɟɬ ɫɭɳɟɫɬɜɟɧɧɵɯ ɢɡɦɟɧɟɧɢɣ ɜ ɞɢɧɚɦɢɤɟ ɢɯ ɜɨɡɪɚɫɬɧɨɝɨ ɪɚɡɜɢɬɢɹ ȼɨ ɜɡɪɨɫɥɵɯ ɫɬɚɞɢɹɯ ɩɚɭɤɨɨɛɪɚɡɧɵɯ ɩɪɨɢɫɯɨɞɹɬ ɚɞɚɩɬɢɜ ɧɵɟ ɢɡɦɟɧɟɧɢɹ ɥɨɜɱɢɯ ɫɟɬɟɣ ɜ ɡɚɜɢɫɢɦɨɫɬɢ ɨɬ ɪɚɡɦɟɪɨɜ ɤɨɥɢɱɟɫɬɜɚ ɥɟɬɚɸɳɢɯ ɧɚɫɟɤɨɦɵɯ ɢ ɜɧɟɲɧɢɯ ɭɫɥɨɜɢɣ ɦɟɫɬɨɨɛɢɬɚɧɢɣ
ȼɥɢɹɧɢɟ ɯɪɨɧɢɱɟɫɤɢɯ ɚɧɬɪɨɩɨɝɟɧɧɵɯ ɮɚɤɬɨɪɨɜ ɧɚ ɫɬɪɭɤɬɭɪɭ ɥɨɜɱɢɯ
ɫɟɬɟɣ Larinioides patagiatus
ȼ ɤɚɱɟɫɬɜɟ ɨɬɜɟɬɧɨɣ ɪɟɚɤɰɢɟɣ ɩɚɭɤɨɜ ɧɚ ɜɨɡɞɟɣɫɬɜɢɟ ɯɪɨɧɢɱɟɫɤɢɯ ɚɧɬɪɨɩɨ ɝɟɧɧɵɯ ɮɚɤɬɨɪɨɜ ɪɚɫɫɦɚɬɪɢɜɚɥɨɫɶ ɢɡɦɟɧɟɧɢɹ ɜ ɩɨɜɟɞɟɧɢɢ ɩɪɢ ɫɬɪɨɢɬɟɥɶɫɬɜɟ ɥɨɜ ɱɢɯ ɫɟɬɟɣ Ⱥɜɬɨɬɪɚɧɫɩɨɪɬ ɧɚɢɛɨɥɟɟ ɪɚɫɩɪɨɫɬɪɚɧɟɧɧɵɣ ɜɢɞ ɚɧɬɪɨɩɨɝɟɧɧɨɝɨ ɡɚ ɝɪɹɡɧɟɧɢɹ ɫɨɜɪɟɦɟɧɧɨɣ ɭɪɛɚɧɢɡɢɪɨɜɚɧɧɨɣ ɫɪɟɞɵ ɉɚɭɤɢ ɚɤɬɢɜɧɨ ɢɫɩɨɥɶɡɭɸɬ ɛɨɪɞɸɪɵ ɚɜɬɨɞɨɪɨɠɧɵɯ ɦɨɫɬɨɜ ɜ ɤɚɱɟɫɬɜɟ ɨɩɨɪɵ ɞɥɹ ɩɥɟɬɟɧɢɹ ɫɜɨɢɯ ɥɨɜɱɢɯ ɫɟ ɬɟɣ ɩɨɷɬɨɦɭ ɫɬɪɭɤɬɭɪɚ ɩɨɫɥɟɞɧɢɯ ɦɨɠɟɬ ɢɫɩɨɥɶɡɨɜɚɬɶɫɹ ɞɥɹ ɨɰɟɧɤɢ ɫɬɟɩɟɧɢ ɜɥɢɹɧɢɹ ɜɵɯɥɨɩɧɵɯ ɝɚɡɨɜ ɜɢɛɪɚɰɢɢ ɢ ɲɭɦɚ ɚɜɬɨɦɨɛɢɥɟɣ ɧɚ ɩɨɜɟɞɟɧɢɟ ɩɚɭɤɨɜ Ⱦɥɹ ɨɰɟɧɤɢ ɜɥɢɹɧɢɹ ɚɜɬɨɬɪɚɧɫɩɨɪɬɚ ɧɚ ɩɚɭɤɨɜ-ɤɪɭɝɨɩɪɹɞɨɜ ɢɫɩɨɥɶɡɨɜɚɥɢɫɶ ɞɜɚ ɭɱɚɫɬɤɚ ɤɨɧɬɪɨɥɶɧɵɣ- ɧɟɞɟɣɫɬɜɭɸɳɢɣ ɦɨɫɬɨɜɨɣ ɩɟɪɟɟɡɞ ɢ ɨɩɵɬɧɵɣ - ɞɟɣɫɬɜɭɸ ɳɢɣ ɦɨɫɬɨɜɨɣ ɩɟɪɟɟɡɞ ɧɚ ɤɨɬɨɪɵɯ ɨɩɢɫɵɜɚɥɢɫɶ ɥɨɜɱɢɟ ɫɟɬɢ ɩɚɭɤɨɜ ȼ ɤɚɱɟɫɬɜɟ ɨɛɴɟɤɬɨɜ ɢɫɫɥɟɞɨɜɧɢɣ ɢɫɩɨɥɶɡɨɜɚɥɢɫɶɩɚɭɤɢ ɜɢɞɚ Larinioides patagiatus. Ʌɨɜɱɚɹ ɫɟɬɶ L. patagiatus ɢɦɟɟɬ ɯɚɪɚɤɬɟɪɧɭɸ ɫɬɪɭɤɬɭɪɭ ɬɢɩɢɱɧɭɸ ɞɥɹ ɞɚɧɧɨɝɨ ɜɢɞɚ ɩɚɭ
ɤɨɜ ɪɢɫ
Ⱦɥɹ ɰɟɧɬɪɚɥɶɧɨɣ ɡɨɧɵ ɥɨɜɱɢɯ ɫɟɬɟɣ ɩɚɭɤɨɜ ɜɢɞɚ L. patagiatus ɯɚɪɚɤɬɟɪɧɵ ɧɟɛɨɥɶɲɢɟ ɪɚɡɦɟɪɵ±ɫɦ ɢ ɤɨɥɢɱɟɫɬɜɨ ɜɢɬɤɨɜ ɪɚɜɧɨɟ Ɋɚɡɦɟɪɵ ɫɜɨɛɨɞ ɧɨɣ ɡɨɧɵ ɜɚɪɶɢɪɭɸɬ ɜ ɩɪɟɞɟɥɚɯ±ɫɦ Ʌɨɜɱɚɹ ɡɨɧɚ ɫɨɫɬɨɹɳɚɹ ɢɡ ɤɥɟɣɤɢɯ ɥɨɜɱɢɯ ɧɢɬɟɣ ɢ ɪɚɞɢɚɥɶɧɨ ɪɚɫɩɨɥɨɠɟɧɧɵɯ ɪɚɞɢɭɫɨɜ ɧɟɡɧɚɱɢɬɟɥɶɧɨ ɜɵɬɹɧɭɬɚ ɜɧɢɡ Ɋɚɫɫɬɨɹɧɢɟ ɦɟɠɞɭ ɥɨɜɱɢɦɢ ɧɢɬɹɦɢ±ɫɦ ɱɢɫɥɨ ɪɚɞɢɭɫɨɜ ɪɚɜɧɨ ɢ
ϲϳ
ɤɨɥɢɱɟɫɬɜɨ ɥɨɜɱɢɯ ɧɢɬɟɣ ɪɚɜɧɨ ± ȼ ɰɟɥɨɦ ɥɨɜɱɚɹ ɡɨɧɚ ɩɚɭɤɨɜ ɜɢɞɚ L. patagiatus L. patagiatus ɞɨɫɬɚɬɨɱɧɨ ɛɨɥɶɲɚɹ ɢ ɯɨɪɨɲɨ ɡɚɦɟɬɧɚɹ ɇɟɨɛɯɨɞɢɦɨ ɨɬ ɦɟɬɢɬɶ ɱɬɨ ɩɪɟɞɫɬɚɜɢɬɟɥɢ ɪɚɫɫɦɚɬɪɢɜɚɟɦɨɝɨ ɜɢɞɚ ɨɯɨɬɹɬɫɹ ɧɚ ɧɚɫɟɤɨɦɵɯ ɜ ɪɟɱ ɧɵɯ ɩɪɨɬɨɤɚɯ ɢ ɪɟɤɚɯ.
ɋɪɚɜɧɢɬɟɥɶɧɵɣ ɚɧɚɥɢɡ ɫɟɞɧɟɫɬɚɬɢɫɬɢɱɟɫɤɢɯ ɩɨɤɚɡɚɬɟɥɟɣ ɥɨɜɱɢɯ ɫɟɬɟɣ L. patagiatus ɬɚɛɥ ɩɨɡɜɨɥɢɥ ɡɚɦɟɬɢɬɶ ɱɬɨ ɩɨ ɜɫɟɦ ɩɨɤɚɡɚɬɟɥɹɦ ɪɚɡɦɟɪɵ ɥɨɜɱɢɯ ɫɟɬɟɣ L. patagiatus ɜ ɤɨɧɬɪɨɥɟ ɩɪɟɜɵɲɚɸɬ ɚɧɚɥɨɝɢɱɧɵɟ ɩɨɤɚɡɚɬɟɥɢ ɧɚ ɨɩɵɬɧɨɦ ɭɱɚɫɬɤɟ
Ɋɢɫ Ʌɨɜɱɚɹ ɫɟɬɶ L. Patagiatus-ɤɨɧɬɪɨɥɶɧɵɣ ɭɱɚɫɬɨɤ
Ⱦɥɹ ɨɩɵɬɧɨɝɨ ɢ ɤɨɧɬɪɨɥɶɧɨɝɨ ɭɱɚɫɬɤɨɜ ɯɚɪɚɤɬɟɪɧɨ ɪɚɡɧɨɟ ɱɢɫɥɨ ɜɢɬɤɨɜ ɜ ɰɟɧɬɪɚɥɶɧɨɣ ɡɨɧɟ ɢ ɫɨɨɬɜɟɬɫɬɜɟɧɧɨ ɱɬɨ ɭɤɚɡɵɜɚɟɬ ɧɚ ɫɭɳɟɫɬɜɟɧɧɨɟ ɜɥɢɹɧɢɟ ɚɜɬɨɬɪɚɧɫɩɨɪɬɚ ɧɚ ɚɥɝɨɪɢɬɦ ɢɧɫɬɢɧɤɬɢɜɧɨɝɨ ɩɨɜɟɞɟɧɢɹ ɩɚɭɤɨɜ ɑɢɫɥɨ ɜɢɬɤɨɜ ɭ ɦɧɨɝɢɯ ɜɢɞɨɜ ɹɜɥɹɟɬɫɹ ɜɢɞɨɜɵɦ ɩɪɢɡɧɚɤɨɦ ɫɬɪɭɤɬɭɪɵ ɥɨɜɱɢɯ ɫɟɬɟɣ Ʌɨɜɱɢɟ ɫɟɬɢ L. patagiatus ɧɚɯɨɞɹɳɢɟɫɹ ɧɚ ɤɨɧɬɪɨɥɶɧɨɦ ɭɱɚɫɬɤɟ ɨɬɥɢɱɚɸɬɫɹ ɛɨɥɶɲɢɦɢ ɪɚɡɦɟ
ϲϴ
ɪɚɦɢ ɥɨɜɱɟɣ ɢ ɫɜɨɛɨɞɧɨɣ ɡɨɧ ɞɥɢɧɨɣ ɢ ɱɢɫɥɨɦ ɪɚɞɢɭɫɨɜ ɤɨɥɢɱɟɫɬɜɨɦ ɥɨɜɱɢɯ ɧɢ ɬɟɣ ɪɚɡɦɟɪɚɦɢ ɰɟɧɬɪɚɥɶɧɨɣ ɡɨɧɵ Ɋɢɫ Ʌɨɜɱɢɟ ɫɟɬɢ ɧɚ ɨɩɵɬɧɨɦ ɭɱɚɫɬɤɟ ɦɟɧɶɲɟ ɩɨ ɪɚɡɦɟɪɚɦ ɜ ɫɪɟɞɧɟɦ ɧɚ ±ɫɦ ɛɨɥɟɟ ɤɨɦɩɚɤɬɧɵɟ Ɋɢɫ Ʌɨɜɱɚɹ ɫɟɬɶ ɧɚɯɨɞɢɬɫɹ ɛɥɢɠɟ ɤ ɬɜɟɪɞɨɣ ɩɨɜɟɪɯɧɨɫɬɢ Ɋɚɫɫɬɨɹɧɢɹ ɦɟɠɞɭ ɥɨɜɱɢɦɢ ɧɢɬɹɦɢ ɜ ɫɟɬɤɚɯ ɩɚɭɤɨɜ ɧɚ ɨɩɵɬɧɵɯ ɢ ɤɨɧɬɪɨɥɶɧɵɯ ɭɱɚɫɬɤɚɯ ɧɟ ɢɡɦɟɧɹɸɬɫɹ ɢ ɪɚɜɧɵ: 0,3±ɫɚɧɬɢɦɟɬɪɚɦ Ⱥɧɚɥɢɡ ɨɫɧɨɜɧɵɯ ɩɨɤɚɡɚɬɟɥɟɣ ɫɬɪɭɤɬɭɪɵ ɥɨɜɱɢɯ ɫɟɬɟɣ ɞɥɢɧ ɪɚ ɞɢɭɫɨɜ ɪɚɡɦɟɪɨɜ ɥɨɜɱɟɣ ɡɨɧɵ ɪɚɡɦɟɪɨɜ ɫɜɨɛɨɞɧɨɣ ɡɨɧɵ ɩɨɤɚɡɚɥ ɱɬɨ ɫɟɬɢ ɢ ɧɚ ɨɩɵɬɧɨɦ ɢ ɤɨɧɬɪɨɥɶɧɨɦ ɭɱɚɫɬɤɚɯ ɜɵɬɹɧɭɬɵ ɜɧɢɡ Ʉɨɷɮɮɢɰɢɟɧɬ ɚɫɢɦɦɟɬɪɢɢ ɞɥɹ ɥɨɜɱɢɯ ɫɟɬɟɣ L. patagia-tus, ɪɚɫɩɨɥɨɠɟɧɧɵɯ ɧɚ ɤɨɧɬɪɨɥɶɧɨɦ ɭɱɚɫɬɤɟ ɪɚɜɟɧ %, ɞɥɹ ɥɨɜɱɢɯ ɫɟɬɟɣ ɧɚ ɨɩɵɬɧɨɦ ɭɱɚɫɬɤɟ 60 %.
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Ɍɚɛɥɢɰɚ |
ɋɪɟɞɧɢɟ ɡɧɚɱɟɧɢɹ ɩɨɤɚɡɚɬɟɥɟɣ ɥɨɜɱɢɯ ɫɟɬɟɣ L. patagiatus |
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ɧɚ ɤɨɧɬɪɨɥɶɧɨɦ ɢ ɨɩɵɬɧɨɦ ɭɱɚɫɬɤɚɯ |
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ɉɨɤɚɡɚɬɟɥɢ ɫɬɪɭɤɬɭɪɵ ɥɨɜɱɟɣ ɫɟɬɢ |
Ɂɧɚɱɟɧɢɹ ɩɨɤɚɡɚɬɟɥɟɣ |
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ɤɨɧɬɪɨɥɶ |
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ɨɩɵɬ |
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ɑɢɫɥɨ ɜɢɬɤɨɜ ɜ ɰɟɧɬɪɚɥɶɧɨɣ ɡɨɧɟ |
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5 |
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4 |
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Ɋɚɡɦɟɪ ɰɟɧɬɪɚɥɶɧɨɣ ɡɨɧɵ |
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ɜɜɟɪɯ |
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ɜɩɪɚɜɨ |
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ɜɧɢɡ |
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ɜɥɟɜɨ |
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Ɋɚɡɦɟɪ ɫɜɨɛɨɞɧɨɣ ɡɨɧɵ |
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ɜɜɟɪɯ |
1 |
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ɜɩɪɚɜɨ |
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ɜɧɢɡ |
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ɜɥɟɜɨ |
1, |
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ɑɢɫɥɨ ɪɚɞɢɭɫɨɜ |
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23,00 |
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* |
Ⱦɥɢɧɚ ɪɚɞɢɭɫɨɜ |
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ɜɜɟɪɯ |
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ɜɩɪɚɜɨ |
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* |
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ɜɧɢɡ |
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ɜɥɟɜɨ |
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11 |
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Ɉɤɨɧɱɚɧɢɟ ɬɚɛɥ |
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ɉɨɤɚɡɚɬɟɥɢ ɫɬɪɭɤɬɭɪɵ ɥɨɜɱɟɣ ɫɟɬɢ |
Ɂɧɚɱɟɧɢɹ ɩɨɤɚɡɚɬɟɥɟɣ |
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ɤɨɧɬɪɨɥɶ |
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ɨɩɵɬ |
Ɋɚɡɦɟɪ ɥɨɜɱɟɣ ɡɨɧɵ |
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ɜɜɟɪɯ |
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ɜɩɪɚɜɨ |
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ɜɧɢɡ |
14 |
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ɜɥɟɜɨ |
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ɑɢɫɥɨ ɥɨɜɱɢɯ ɧɢɬɟɣ |
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ɜɜɟɪɯ |
31 |
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19 * |
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ɜɩɪɚɜɨ |
30 |
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* |
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ɜɧɢɡ |
33 3 |
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* |
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ɜɥɟɜɨ |
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22 |
Ɋɚɫɫɬɨɹɧɢɟ ɦɟɠɞɭ ɥɨɜɱɢ |
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ɜɜɟɪɯ |
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ɦɢ ɧɢɬɹɦɢ |
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ɜɩɪɚɜɨ |
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ɜɧɢɡ |
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ɜɥɟɜɨ |
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Ɋɚɫɫɬɨɹɧɢɟ ɨɬ ɫɟɪɟɞɢɧɵ ɰɟɧɬɪɚɥɶɧɨɣ ɡɨ |
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90 |
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ϲϵ
ɧɵ ɥɨɜɱɟɣ ɫɟɬɢ ɞɨ ɡɟɦɥɢ
ɉɪɢɦɟɱɚɧɢɟ Ɋɚɡɦɟɪɵ ɞɥɢɧɵ ɢ ɪɚɫɫɬɨɹɧɢɹ ɩɪɢɜɟɞɟɧɵ ɜ ɫɚɧɬɢɦɟɬɪɚɯ
ɉɚɭɤɢ ɩɢɬɚɸɬɫɹ ɧɚɫɟɤɨɦɵɦɢ ɢ ɫɩɨɫɨɛɧɵ ɧɚɤɚɩɥɢɜɚɬɶ ɜ ɫɜɨɟɦ ɬɟɥɟ ɬɨɤɫɢɱɟ ɫɤɢɟ ɜɟɳɟɫɬɜɚ ɤɨɬɨɪɵɟ ɢɡɦɟɧɹɸɬ ɢɯ ɩɨɜɟɞɟɧɢɟ ɱɬɨ ɨɬɪɚɠɚɟɬɫɹ ɜ ɫɬɪɭɤɬɭɪɟ ɥɨɜ ɱɢɯ ɫɟɬɟɣ ɂɡɦɟɧɟɧɢɹ ɜ ɩɨɜɟɞɟɧɢɢ ɠɢɜɨɬɧɵɯ ɮɢɤɫɢɪɭɸɬɫɹ ɜ ɫɬɪɭɤɬɭɪɟ ɥɨɜɱɢɯ ɫɟɬɟɣ ȼ ɥɨɜɱɢɯ ɫɟɬɹɯ L. patagiatus ɧɚ ɨɩɵɬɧɨɦ ɭɱɚɫɬɤɟ ɨɬɦɟɱɟɧɨ ɬɢɩɨɜ ɧɚɪɭɲɟ ɧɢɣ ɫɬɪɭɤɬɭɪɵ ɢɥɢ ɚɧɨɦɚɥɶɧɵɯ ɨɬɤɥɨɧɟɧɢɣ ɫɛɥɢɠɟɧɢɟ ɪɚɞɢɭɫɨɜ ɪɚɫɳɟɩɥɟɧɢɟ ɪɚɞɢɭɫɨɜ ɩɟɪɟɤɪɟɫɬ ɪɚɞɢɭɫɨɜ ɧɨɠɧɢɰɵ ɢɫɤɪɢɜɥɟɧɢɟ ɪɚɞɢɭɫɨɜ ɧɚɥɢɱɢɟ ɥɢɲɧɟɣ ɥɨɜɱɟɣ ɧɢɬɢ ɜ ɫɟɤɬɨɪɟ ɬɚɛɥ ɤɨɬɨɪɵɟ ɧɟ ɜɫɬɪɟɱɚɸɬɫɹ ɜ ɤɨɧɬɪɨɥɶɧɵɯ ɝɪɭɩɩɚɯ ɡɚ ɢɫɤɥɸɱɟɧɢɟɦ ɚɧɨɦɚɥɢɢ ɬɢɩɚ©ɪɚɫɳɟɩɥɟɧɢɟ ɪɚɞɢɭɫɨɜªɇɚɢɛɨɥɟɟ ɱɚɫɬɵɦɢ ɨɬ ɤɥɨɧɟɧɢɹɦɢ ɜ ɨɩɵɬɟ ɹɜɥɹɸɬɫɹ ɧɚɥɢɱɢɟ ɥɢɲɧɟɣ ɥɨɜɱɟɣ ɧɢɬɢ ɜ ɫɟɤɬɨɪɟ ɢ ɫɛɥɢɠɟ ɧɢɟ ɪɚɞɢɭɫɨɜ ɉɨɫɥɟɞɧɟɟ ɤɚɤ ɩɪɚɜɢɥɨ ɧɚɪɭɲɚɟɬ ɨɛɳɭɸ ɫɬɪɭɤɬɭɪɭ ɥɨɜɱɟɣ ɫɟɬɢ
ϳϬ
Ɋɢɫ Ʌɨɜɱɚɹ ɫɟɬɶ L. patagiatus ɨɩɵɬɧɵɣ ɭɱɚɫɬɨɤ
ϳϭ
Ɍɚɛɥɢɰɚ
Ⱥɧɨɦɚɥɢɢ ɜ ɫɟɬɹɯ ɩɨɥɨɜɨɡɪɟɥɵɯ ɫɚɦɨɤ L. patagiatus ɧɚ ɤɨɧɬɪɨɥɶɧɨɦ ɢ ɨɩɵɬɧɨɦ ɭɱɚɫɬɤɚɯ
ɉɨɤɚɡɚɬɟɥɢ ɫɬɪɭɤɬɭɪɵ ɥɨɜɱɟɣ |
Ɉɩɵɬ |
Ʉɨɧɬɪɨɥɶ |
ɫɟɬɢ |
|
|
|
|
|
|
|
|
ɋɛɥɢɠɟɧɢɟ ɪɚɞɢɭɫɨɜ |
4 1 |
0 |
|
|
|
Ɋɚɫɳɟɩɥɟɧɢɟ ɪɚɞɢɭɫɨɜ |
2 1 |
2 1 |
|
|
|
©ɇɨɠɧɢɰɵª |
3 1 |
0 |
ɂɫɤɪɢɜɥɟɧɢɟ ɪɚɞɢɭɫɨɜ |
3 1 |
0 |
Ʌɢɲɧɹɹ ɧɢɬɶ ɜ ɫɟɤɬɨɪɟ |
4 1 |
0 |
ɋɥɟɞɨɜɚɬɟɥɶɧɨ ɧɚ ɩɟɪɜɵɯ ɫɬɚɞɢɹɯ ɯɪɨɧɢɱɟɫɤɢ ɞɟɣɫɬɜɭɸɳɟɝɨ ɧɟɝɚɬɢɜɧɨɝɨ ɮɚɤɬɨɪɚ ɭɜɟɥɢɱɢɜɚɟɬɫɹ ɤɨɥɢɱɟɫɬɜɨ ɧɚɪɭɲɟɧɢɣ ɜ ɩɨɜɟɞɟɧɱɟɫɤɢɯ ɚɥɝɨɪɢɬɦɚɯ ɩɚɭ ɤɨɜ ɤɨɬɨɪɵɟ ɦɨɝɭɬ ɩɪɢɜɨɞɢɬɶ ɤ ɫɬɪɭɤɬɪɧɵɦ ɢɡɦɟɧɟɧɢɹɦ ɜ ɢɯ ɥɨɜɱɢɯ ɡɨɧɚɯ
ɋɬɚɬɢɫɬɢɱɟɫɤɢɣ ɮɚɤɬɨɪɧɵɣ ɚɧɚɥɢɡ ɫ ɢɫɩɨɥɶɡɨɜɚɧɢɟɦ ɦɟɬɨɞɚ ɝɥɚɜɧɵɯ ɤɨɦ ɩɨɧɟɧɬ ɩɨɡɜɨɥɢɥ ɜɵɞɟɥɢɬɶ ɨɫɧɨɜɧɵɯ ɧɟɡɚɜɢɫɢɦɵɯ ɮɚɤɬɨɪɚ ɨɩɪɟɞɟɥɹɸɳɢɯ ɫɬɪɭɤɬɭɪɭ ɥɨɜɱɢɯ ɫɟɬɟɣ ȼ ɤɨɧɬɪɨɥɶɧɨɣ ɝɪɭɩɩɟ ɩɨɤɚɡɚɬɟɥɟɣ ɥɨɜɱɢɯ ɫɟɬɟɣ ɜɵɹɜ ɥɟɧɧɨ ɨɛɳɟɣ ɞɢɫɩɟɪɫɢɢ ɜ ɨɩɵɬɧɨɣ ɝɪɭɩɩɟ- 72 % ɱɬɨ ɭɤɚɡɵɜɚɟɬ ɧɚ ɦɟɧɶɲɭɸ ɫɬɪɭɤɬɭɪɢɪɨɜɚɧɧɨɫɬɶ ɥɨɜɱɢɯ ɫɟɬɟɣ ɩɚɭɤɨɜ ɜ ɨɩɵɬɟ ɬɚɛɥ
Ɍɚɛɥɢɰɚ Ɋɚɫɩɪɟɞɟɥɟɧɢɟ ɨɛɳɟɣ ɞɢɫɩɟɪɫɢɢ ɩɨɤɚɡɚɬɟɥɟɣ ɩɨ ɝɥɚɜɧɵɦ ɤɨɦɩɨɧɟɧɬɚɦ
ɥɨɜɱɢɯ ɫɟɬɟɣ L. patagiatus ɧɚ ɨɩɵɬɧɨɦ ɢ ɤɨɧɬɪɨɥɶɧɨɦ ɭɱɚɫɬɤɚɯ
Ʉɨɦɩɨɧɟɧɬɵ |
Ɉɛɳɚɹ ɞɢɫɩɟɪɫɢɹ |
||
Ʉɨɧɬɪɨɥɶɧɵɣ ɭɱɚɫɬɨɤ |
Ɉɩɵɬɧɵɣ ɭɱɚɫɬɨɤ |
||
|
|||
1 |
36,789 |
22,092 |
|
2 |
22,349 |
14,534 |
|
3 |
11,566 |
13,237 |
|
4 |
6,628 |
8,849 |
|
ɋɪɚɜɧɢɬɟɥɶɧɵɣ ɚɧɚɥɢɡ ɞɢɫɩɟɪɫɢɣ ɝɥɚɜɧɵɯ ɤɨɦɩɨɧɟɧɬ ɜ ɤɨɧɬɪɨɥɶɧɵɯ ɢ ɨɩɵɬɧɵɯ ɜɵɛɨɪɤɚɯ ɜɵɹɜɢɥ ɫɧɢɠɟɧɢɟ ɤɨɥɢɱɟɫɬɜɚ ɜɡɚɢɦɨɫɜɹɡɚɧɧɵɯ ɩɨɤɚɡɚɬɟɥɟɣ ɜ ɩɟɪɜɨɣ-ɜɢɞɨɜɨɣ ɢ ɜɬɨɪɨɣ ɩɢɳɟɜɨɣ ɤɨɦɩɨɧɟɧɬɟ ɩɪɢ ɞɟɣɫɬɜɢɢ ɚɧɬɪɨɩɨɝɟɧɧɵɯ ɮɚɤɬɨɪɨɜ ȼ ɬɪɟɬɶɟɣ ɢ ɱɟɬɜɺɪɬɨɣ ɛɢɨɬɨɩɢɱɟɫɤɢɯ ɤɨɦɩɨɧɟɧɬɚɯ ɩɪɨɢɫɯɨɞɢɬ ɭɜɟɥɢ ɱɟɧɢɟ ɤɨɥɢɱɟɫɬɜɚ ɤɨɪɪɟɥɹɰɢɨɧɧɵɯ ɜɡɚɢɦɨɞɟɣɫɬɜɢɣ ɬɚɛɥ ɂɧɬɟɪɩɪɟɬɢɪɭɹ ɜɵɞɟɥɟɧɧɵɟ ɤɨɦɩɨɧɟɧɬɵ ɤɨɧɬɪɨɥɶɧɵɯ ɫɟɬɟɣ L. patagiatus ɫɥɟɞɭɟɬ ɨɬɦɟɬɢɬɶ ɱɬɨ ɜ ɩɟɪɜɨɣ ɜɢɞɨɜɨɣ ɧɚɫɥɟɞɫɬɜɟɧɧɨɣ ɤɨɦɩɨɧɟɧɬɟ ɜɤɥɸɱɚɸɳɟɣ ɫɬɪɭɤɬɭɪɭ ɥɨɜɱɟɣ ɫɟ ɬɢ ɫɨɫɪɟɞɨɬɨɱɟɧɨ ɨɛɳɟɣ ɞɢɫɩɟɪɫɢɢ ɉɟɪɜɚɹ ɤɨɦɩɨɧɟɧɬɚ ɨɛɴɟɞɢɧɹɟɬ ɩɨɤɚɡɚ ɬɟɥɢ ɥɨɜɱɟɣ ɡɨɧɵ ɞɥɢɧɵ ɪɚɞɢɭɫɨɜ ɪɚɡɦɟɪɵ ɥɨɜɱɟɣ ɡɨɧɵ ȼɬɨɪɚɹ ɤɨɦɩɨɧɟɧɬɚ ɩɢɳɟɜɚɹ ɨɩɢɫɵɜɚɟɬ ɜɡɚɢɦɨɫɜɹɡɶ ɦɟɠɞɭ ɤɨɥɢɱɟɫɬɜɨɦ ɥɨɜɱɢɯ ɧɢɬɟɣ ɜ ɫɟɤɬɨɪɟ ɫ ɱɢɫɥɨɦ ɪɚɞɢɭɫɨɜ ɪɚɡɦɟɪɚɦɢ ɰɟɧɬɪɚɥɶɧɨɣ ɢ ɫɜɨɛɨɞɧɨɣ ɡɨɧ Ɉɩɪɟɞɟɥɹɸɳɢɦ ɜɨ ɜɬɨɪɨɣ ɤɨɦɩɨɧɟɧɬɟ ɹɜɥɹɟɬɫɹ ɤɨɥɢɱɟɫɬɜɨ ɥɨɜɱɢɯ ɧɢɬɟɣ ɤɨ ɬɨɪɵɟ ɮɨɪɦɢɪɭɸɬ ɥɨɜɱɭɸ ɡɨɧɭ ɧɟɡɚɜɢɫɢɦɨ ɨɬ ɞɥɢɧ ɪɚɞɢɭɫɨɜ ɢ ɪɚɫɫɬɨɹɧɢɣ ɦɟɠɞɭ ɥɨɜɱɢɦɢ ɧɢɬɹɦɢ ɢ ɯɨɪɨɲɨ ɤɨɪɪɟɥɢɪɭɟɬ ɫ ɩɨɤɚɡɚɬɟɥɹɦɢ ɰɟɧɬɪɚɥɶɧɨɣ ɢ ɫɜɨɛɨɞɧɨɣ ɡɨɧɵ Ɍɪɟɬɶɹ ɢ ɱɟɬɜɺɪɬɚɹ ɛɢɨɬɨɩɢɱɟɫɤɢɟ ɤɨɦɩɨɧɟɧɬɵ ɨɛɴɟɞɢɧɹɸɬ ɧɢɠɧɢɟ ɢ ɛɨɤɨ ɜɵɟ ɩɨɤɚɡɚɬɟɥɢ ɚɫɫɢɦɟɬɪɢɢ ɪɚɡɦɟɪɨɜ ɰɟɧɬɪɚɥɶɧɨɣ ɫɜɨɛɨɞɧɨɣ ɡɨɧ ɤɨɥɢɱɟɫɬɜɨ ɪɚ ɞɢɭɫɨɜ ɛɨɤɨɜɵɟ ɢ ɜɟɪɯɧɢɟ ɩɨɤɚɡɚɬɟɥɢ ɤɨɥɢɱɟɫɬɜɚ ɥɨɜɱɢɯ ɧɢɬɟɣ
ϳϮ
ɋɬɪɭɤɬɭɪɚ ɝɥɚɜɧɵɯ ɤɨɦɩɨɧɟɧɬ |
|
Ɍɚɛɥɢɰɚ |
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ɜ ɥɨɜɱɢɯ ɫɟɬɹɯ L. patagiatus |
ɤɨɧɬɪɨɥɶɧɨɣ ɜɵɛɨɪɤɢ |
|
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ɉɨɤɚɡɚɬɟɥɢ |
Ɏɚɤɬɨɪ |
|
Ɏɚɤɬɨɪ |
Ɏɚɤɬɨɪ |
Ɏɚɤɬɨɪ |
ɑɢɫɥɨ ɜɢɬɤɨɜ ɜ ɰɟɧɬɪɚɥɶɧɨɣ |
0,132 |
|
0,588* |
0,069 |
0,405* |
ɡɨɧɟ |
|
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|
|
|
|
|
Ɋɚɡɦɟɪ ɰɟɧɬɪɚɥɶɧɨɣ ɡɨɧɵ |
0,210 |
|
0,795* |
0,481* |
0,038 |
ɜɜɟɪɯ |
|
||||
|
|
|
|
|
|
Ɋɚɡɦɟɪ ɰɟɧɬɪɚɥɶɧɨɣ ɡɨɧɵ |
0,180 |
|
0,709* |
0,479* |
0,169 |
ɜɩɪɚɜɨ |
|
||||
|
|
|
|
|
|
Ɋɚɡɦɟɪ ɰɟɧɬɪɚɥɶɧɨɣ ɡɨɧɵ ɜɧɢɡ |
0,119 |
|
0,783* |
0,564* |
0,118 |
Ɋɚɡɦɟɪ ɰɟɧɬɪɚɥɶɧɨɣ ɡɨɧɵ ɜɥɟ |
0,002 |
|
0,787* |
0,568* |
0,008 |
ɜɨ |
|
||||
|
|
|
|
|
|
Ɋɚɡɦɟɪ ɫɜɨɛɨɞɧɨɣ ɡɨɧɵ |
0,119 |
|
0,422* |
0,307 |
0,269 |
ɜɜɟɪɯ |
|
||||
|
|
|
|
|
|
Ɋɚɡɦɟɪ ɫɜɨɛɨɞɧɨɣ ɡɨɧɵ |
0,436* |
|
0,547* |
0,319 |
0,366 |
ɜɩɪɚɜɨ |
|
||||
|
|
|
|
|
|
Ɋɚɡɦɟɪ ɫɜɨɛɨɞɧɨɣ ɡɨɧɵ ɜɧɢɡ |
0,645* |
|
0,129 |
0,449* |
0,282 |
Ɋɚɡɦɟɪ ɫɜɨɛɨɞɧɨɣ ɡɨɧɵ |
0,372 |
|
0,634* |
0,449* |
0,345 |
ɜɥɟɜɨ |
|
||||
|
|
|
|
|
|
ɑɢɫɥɨ ɪɚɞɢɭɫɨɜ |
0,196 |
|
0,521* |
0,402* |
0,378 |
Ⱦɥɢɧɚ ɪɚɞɢɭɫɨɜ ɜɜɟɪɯ |
0,789* |
|
0,324 |
0,267 |
0,095 |
Ⱦɥɢɧɚ ɪɚɞɢɭɫɨɜ ɜɩɪɚɜɨ |
0,810* |
|
0,079 |
0,078 |
0,279 |
Ⱦɥɢɧɚ ɪɚɞɢɭɫɨɜ ɜɧɢɡ |
0,854* |
|
0,009 |
0,215 |
0,390 |
Ⱦɥɢɧɚ ɪɚɞɢɭɫɨɜ ɜɥɟɜɨ |
0,892* |
|
0,094 |
0,195 |
0,126 |
Ɋɚɡɦɟɪ ɥɨɜɱɟɣ ɡɨɧɵ ɜɜɟɪɯ |
0,769* |
|
0,409* |
0,273 |
0,149 |
Ɋɚɡɦɟɪ ɥɨɜɱɟɣ ɡɨɧɵ ɜɩɪɚɜɨ |
0,809* |
|
0,021 |
0,126 |
0,319 |
Ɋɚɡɦɟɪ ɥɨɜɱɟɣ ɡɨɧɵ ɜɧɢɡ |
0,887* |
|
0,122 |
0,224 |
0,233 |
Ɋɚɡɦɟɪ ɥɨɜɱɟɣ ɡɨɧɵ ɜɥɟɜɨ |
0,881* |
|
0,187 |
0,032 |
0,094 |
ɑɢɫɥɨ ɥɨɜɱɢɯ ɧɢɬɟɣ ɜɜɟɪɯ |
0,527* |
|
0,628* |
0,480* |
0,038 |
ɑɢɫɥɨ ɥɨɜɱɢɯ ɧɢɬɟɣ ɜɩɪɚɜɨ |
0,425* |
|
0,489* |
0,591* |
0,136 |
ɑɢɫɥɨ ɥɨɜɱɢɯ ɧɢɬɟɣ ɜɧɢɡ |
0,426* |
|
0,647* |
0,266 |
0,359 |
ɑɢɫɥɨ ɥɨɜɱɢɯ ɧɢɬɟɣ ɜɥɟɜɨ |
0,493* |
|
0,592* |
0,469* |
0,181 |
Ɋɚɫɫɬɨɹɧɢɟ ɦɟɠɞɭ ɥɨɜɱɢɦɢ ɧɢ |
0,764* |
|
0,153 |
0,046 |
0,244 |
ɬɹɦɢ ɜɜɟɪɯ |
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ɉɨɤɚɡɚɬɟɥɢ |
Ɏɚɤɬɨɪ |
|
Ɏɚɤɬɨɪ |
Ɏɚɤɬɨɪ |
Ɏɚɤɬɨɪ |
Ɋɚɫɫɬɨɹɧɢɟ ɦɟɠɞɭ ɥɨɜɱɢɦɢ ɧɢ |
0,751* |
|
0,179 |
0,166 |
0,431* |
ɬɹɦɢ ɜɩɪɚɜɨ |
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Ɋɚɫɫɬɨɹɧɢɟ ɦɟɠɞɭ ɥɨɜɱɢɦɢ ɧɢ |
0,760* |
|
0,455* |
0,113 |
0,040 |
ɬɹɦɢ ɜɧɢɡ |
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Ɋɚɫɫɬɨɹɧɢɟ ɦɟɠɞɭ ɥɨɜɱɢɦɢ ɧɢ |
0,507* |
|
0,384 |
0,059 |
0,395 |
ɬɹɦɢ ɜɥɟɜɨ |
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Ɋɚɫɫɬɨɹɧɢɟ ɨɬ ɫɟɪɟɞɢɧɵ ɰɟɧ |
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ɬɪɚɥɶɧɨɣ ɡɨɧɵ ɥɨɜɱɟɣ ɫɟɬɢ ɞɨ |
0,743* |
|
0,047 |
0,189 |
0,083 |
ɡɟɦɥɢ |
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ϳϯ |
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