Адаптация животных к хроническим факторам
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Ɋɢɫ Ʌɨɜɱɚɹ ɫɟɬɶ L. patagiatus
Ϯϴ
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Ɍɚɛɥɢɰɚ |
ɋɪɟɞɧɢɟ ɡɧɚɱɟɧɢɹ ɩɨɤɚɡɚɬɟɥɟɣ ɥɨɜɱɢɯ ɫɟɬɟɣ L. patagiatus |
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ɉɨɤɚɡɚɬɟɥɢ ɫɬɪɭɤɬɭɪɵ ɥɨɜɱɟɣ ɫɟɬɢ |
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Ɋɚɡɦɟɪɵ ɩɨɤɚɡɚɬɟɥɟɣ ɫɦ |
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ɑɢɫɥɨ ɜɢɬɤɨɜ ɜ ɰɟɧɬɪɚɥɶɧɨɣ ɡɨɧɟ |
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Ɋɚɡɦɟɪ ɰɟɧɬɪɚɥɶɧɨɣ ɡɨɧɵ |
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ɜɜɟɪɯ |
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ɜɩɪɚɜɨ |
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ɜɧɢɡ |
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ɜɥɟɜɨ |
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Ɋɚɡɦɟɪ ɫɜɨɛɨɞɧɨɣ ɡɨɧɵ |
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ɜɜɟɪɯ |
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ɜɩɪɚɜɨ |
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ɜɧɢɡ |
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ɜɥɟɜɨ |
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ɑɢɫɥɨ ɪɚɞɢɭɫɨɜ |
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Ⱦɥɢɧɚ ɪɚɞɢɭɫɨɜ |
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ɜɜɟɪɯ |
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ɜɩɪɚɜɨ |
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ɜɧɢɡ |
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ɜɥɟɜɨ |
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Ɋɚɡɦɟɪ ɥɨɜɱɟɣ ɡɨɧɵ |
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ɜɜɟɪɯ |
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ɜɩɪɚɜɨ |
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ɜɧɢɡ |
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ɜɥɟɜɨ |
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Ʉɨɥɢɱɟɫɬɜɨ ɥɨɜɱɢɯ ɧɢɬɟɣ |
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ɜɜɟɪɯ |
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ɜɩɪɚɜɨ |
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ɜɧɢɡ |
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ɜɥɟɜɨ |
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Ɋɚɫɫɬɨɹɧɢɟ ɦɟɠɞɭ ɥɨɜɱɢɦɢ ɧɢɬɹ |
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ɜɜɟɪɯ |
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ɦɢ |
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ɜɩɪɚɜɨ |
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ɜɧɢɡ |
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ɜɥɟɜɨ |
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Ɋɚɫɫɬɨɹɧɢɟ ɨɬ ɫɟɪɟɞɢɧɵ ɰɟɧɬɪɚɥɶɧɨɣ |
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ɡɨɧɵ ɥɨɜɱɟɣ ɫɟɬɢ ɞɨ ɡɟɦɥɢ |
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ɍɝɨɥ ɧɚɤɥɨɧɚ ɫɟɬɢ ɩɨ ɨɬɧɨɲɟɧɢɸ |
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ɤ ɝɨɪɢɡɨɧɬɚɥɶɧɨɣ ɩɨɜɟɪɯɧɨɫɬɢ |
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Ⱥɧɚɥɢɡ ɝɥɚɜɧɵɯ ɤɨɦɩɨɧɟɧɬ ɜɵɹɜɢɥ ɪɚɫɩɪɟɞɟɥɟɧɢɟ ɞɢɫɩɟɪɫɢɢ ɜ ɩɨɤɚɡɚɬɟɥɹɯ ɥɨɜɱɢɯ ɫɟɬɟɣ ɩɚɭɤɨɜ ɜɢɞɚ L. patagiatus ɨɛɳɟɣ ɞɢɫɩɟɪɫɢɢ ɫɨɫɪɟɞɨɬɨɱɟɧɨ ɜ ɩɟɪɜɵɯ ɤɨɦɩɨɧɟɧɬɚɯ ɹɜɥɹɸɬɫɹ ɫɥɭɱɚɣɧɵɦɢ ɬɚɛɥ
Ϯϵ
ɤɚɡɚɬɟɥɹɦɢ ɥɨɜɱɟɣ ɡɨɧɵ ɱɢɫɥɨɦ ɥɨɜɱɢɯ ɧɢɬɟɣ ɪɚɫɫɬɨɹɧɢɹɦɢ ɦɟɠɞɭ ɥɨɜɱɢɦɢ ɧɢ ɬɹɦɢ ɪɚɡɦɟɪɚɦɢ ɥɨɜɱɟɣ ɡɨɧɵ ɢ ɨɩɪɟɞɟɥɹɸɬ ɪɚɡɦɟɪɵ ɢ ɤɨɦɩɚɤɬɧɨɫɬɶ ɥɨɜɱɟɣ ɫɟɬɢ ȼɬɨɪɚɹ ɤɨɦɩɨɧɟɧɬɚ ɩɢɳɟɜɚɹ ɨɩɢɫɵɜɚɟɬ ɜɡɚɢɦɨɫɜɹɡɶ ɦɟɠɞɭ ɤɨɥɢɱɟɫɬɜɨɦ ɥɨɜɱɢɯ ɧɢɬɟɣ ɜ ɫɟɤɬɨɪɟ ɱɢɫɥɨɦ ɪɚɞɢɭɫɨɜ ɪɚɡɦɟɪɚɦɢ ɰɟɧɬɪɚɥɶɧɵɯ ɢ ɫɜɨɛɨɞɧɵɯ ɡɨɧ Ɉɩ ɪɟɞɟɥɹɸɳɢɦ ɜɨ ɜɬɨɪɨɣ ɤɨɦɩɨɧɟɧɬɟ ɹɜɥɹɟɬɫɹ ɤɨɥɢɱɟɫɬɜɨ ɥɨɜɱɢɯ ɧɢɬɟɣ ɤɨɬɨɪɵɟ ɮɨɪɦɢɪɭɸɬ ɥɨɜɱɭɸ ɡɨɧɭ ɧɟɡɚɜɢɫɢɦɨ ɨɬ ɞɥɢɧ ɪɚɞɢɭɫɨɜ ɢ ɪɚɫɫɬɨɹɧɢɣ ɦɟɠɞɭ ɥɨɜ ɱɢɦɢ ɧɢɬɹɦɢ Ɍɪɟɬɶɹ ɢ ɱɟɬɜɺɪɬɵɟ ɛɢɨɬɨɩɢɱɟɫɤɢɟ ɤɨɦɩɨɧɟɧɬɵ ɨɛɴɟɞɢɧɹɸɬ ɩɨɤɚ ɡɚɬɟɥɢ ɪɚɡɦɟɪɨɜ ɰɟɧɬɪɚɥɶɧɨɣ ɜɟɪɯɧɢɯ ɢ ɛɨɤɨɜɵɯ ɩɨɤɚɡɚɬɟɥɟɣ ɤɨɥɢɱɟɫɬɜ ɥɨɜɱɢɯ ɧɢɬɟɣ ɜ ɫɟɤɬɨɪɟ ɪɚɡɦɟɪɨɜ ɥɨɜɱɟɣ ɡɨɧɵ ɫɜɨɛɨɞɧɵ ɡɨɧ ɢ ɱɢɫɥɚ ɪɚɞɢɭɫɨɜ ɋɥɟɞɨ ɜɚɬɟɥɶɧɨ ɜ ɩɨɜɟɞɟɧɢɢ ɢɫɫɥɟɞɭɟɦɨɝɨ ɜɢɞɚ ɩɚɭɤɨɜ ɫɨɜɨɤɭɩɧɨɫɬɶ ɩɨɤɚɡɚɬɟɥɟɣ ɷɥɟ ɦɟɧɬɨɜ ɩɨɜɟɞɟɧɢɹ ɛɢɨɬɨɩɢɱɟɫɤɨɣ ɫɨɫɬɚɜɥɹɸɳɟɣ ɧɚɢɛɨɥɟɟ ɥɚɛɢɥɶɧɵ ɢ ɡɚɜɢɫɹɬ ɨɬ ɜɥɢɹɧɢɹ ɜɧɟɲɧɢɯ ɮɚɤɬɨɪɨɜ ɨɤɪɭɠɚɸɳɟɣ ɫɪɟɞɵ
ɋɬɪɭɤɬɭɪɚ ɥɨɜɱɢɯ ɫɟɬɟɣ Araneus marmoreus
ɉɚɭɤɢ ɜɢɞɚ A. marmoreus (Clerck, 1758) ɨɬɧɨɫɹɬɫɹ ɤ ɫɚɦɨɦɭ ɨɛɲɢɪɧɨɦɭ ɪɨɞɭ ɤɪɭɝɨɩɪɹɞɨɜ Araneus (Clerck, 1758), ɧɚɫɱɢɬɵɜɚɸɳɟɦɭ ɜɢɞ ɧɚ ɬɟɪɪɢɬɨɪɢɢ Ɋɨɫ ɫɢɢ Ɇɢɯɚɣɥɨɜ ȼ Ɂɚɩɚɞɧɨɣ ɋɢɛɢɪɢ ɨɛɢɬɚɟɬ ɩɨɪɹɞɤɚ ɜɢɞɨɜ ȼ Ɍɨɦɫɤɨɣ ɨɛɥɚɫɬɢ ɨɛɧɚɪɭɠɟɧɨ ɜɢɞɨɜ Ʌɭɤɶɹɧɰɟɜ A. marmoreus, ɤɚɤ ɢ Ⱥ diadematus - ɨɞɧɢ ɢɡ ɧɚɢɛɨɥɟɟ ɯɨɪɨɲɨ ɢɡɭɱɟɧɧɵɯ ɜɢɞɨɜ ɜ ɫɟɦɟɣɫɬɜɟ Araneidae ɉɨɜɫɟɦɟɫɬɧɚɹ ɪɚɫɩɪɨɫɬɪɚɧɟɧɧɨɫɬɶ ɢ ɯɨɪɨɲɨ ɡɚɦɟɬɧɵɟ ɫɟɬɢ ɡɧɚɱɢɬɟɥɶɧɨ ɨɛɥɟɝɱɚɸɬ ɢɡɭɱɟɧɢɟ ɞɚɧɧɨɝɨ ɜɢɞɚ ɜ ɩɨɥɟɜɵɯ ɭɫɥɨɜɢɹɯ
ɋɚɦɤɢ ɩɚɭɤɨɜ A. marmoreus ɹɜɥɹɸɬɫɹ ɤɥɚɫɫɢɱɟɫɤɢɦ ɨɛɴɟɤɬɨɦ ɢɫɫɥɟɞɨɜɚɧɢɣ ɥɨɜɱɢɯ ɫɟɬɟɣ ɤɪɭɝɨɩɪɹɞɨɜ ɉɟɪɜɵɟ ɩɨɥɨɜɨɡɪɟɥɵɟ ɨɫɨɛɢ ɩɨɹɜɥɹɸɬɫɹ ɫɨ ɜɬɨɪɨɣ ɩɨ ɥɨɜɢɧɵ ɢɸɥɹ ɉɥɟɬɭɬ ɫɟɬɢ ɜ ɪɚɧɧɢɟ ɭɬɪɟɧɧɢɟ ɱɚɫɵ ɇɟɩɨɥɨɜɨɡɪɟɥɵɟ ɩɚɭɤɢ ɪɚɫɩɨ ɥɚɝɚɸɬɫɹ ɜ ɰɟɧɬɪɟ ɫɟɬɢ ɜ ɬɨ ɜɪɟɦɹ ɤɚɤ ɩɨɥɨɜɨɡɪɟɥɵɟ ɫɚɦɤɢ ɞɟɥɚɸɬ ɭɛɟɠɢɳɚ ɜ ɜɢ ɞɟ ɫɜɟɪɧɭɬɵɯ ɥɢɫɬɶɟɜ ɞɟɪɟɜɶɟɜ ɫɜɹɡɚɧɧɵɯ ɫ ɰɟɧɬɪɨɦ ɫɢɝɧɚɥɶɧɨɣ ɧɢɬɶɸ Ɋɚɡɦɟɪɵ ɬɟɥɚ ɩɚɭɤɚ ɫɨɫɬɚɜɥɹɸɬ ɜ ɞɥɢɧɭ ± ɦɦ ɋɚɦɤɢ (12±15 ɦɦ ɡɧɚɱɢɬɟɥɶɧɨ ɤɪɭɩɧɟɟ ɫɚɦɰɨɜ ± ɦɦ ɯɚɪɚɤɬɟɪɢɡɭɸɬɫɹ ɛɨɥɟɟ ɞɥɢɧɧɵɦ ɠɢɡɧɟɧɧɵɦ ɰɢɤɥɨɦ ɢ ɩɥɟɬɟɧɢɟɦ ɥɨɜɱɟɣ ɫɟɬɢ ɜ ɬɟɱɟɧɢɟ ɜɫɟɣ ɠɢɡɧɢ Ɉɫɨɛɟɧɧɨɫɬɶɸ ɩɚɭɤɨɜ ɜɢɞɚ A. marmoreus ɹɜɥɹɟɬɫɹ ɰɜɟɬɨɜɨɣ ɩɨɥɢɦɨɪɮɢɡɦ ȼɵɞɟɥɹɸɬ ɞɜɟ ɨɫɧɨɜɧɵɟ ɦɨɪɮɵ ɪɢɫ
ϯϭ
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Ɋɢɫ ɐɜɟɬɨɜɨɣ ɩɨɥɢɦɨɪɮɢɡɦ A. marmoreus: |
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ɚ var marmoreus; ɛ |
var pyramidatus |
ɉɚɭɤɢ A. marmoreus (var marmoreus), ɨɛɢɬɚɸɬ ɩɪɟɢɦɭɳɟɫɬɜɟɧɧɨ ɧɚ ɥɢɫɬɜɟ ɧɢɱɧɵɯ ɩɨɪɨɞɚɯ ɞɟɪɟɜɶɟɜ Ɉɤɪɚɫɤɚ ɛɪɸɲɤɚ ɹɪɤɚɹ ɠɟɥɬɨɝɨ ɢɥɢ ɨɪɚɧɠɟɜɨɝɨ ɰɜɟɬɚ ɫ ɤɪɟɫɬɨɨɛɪɚɡɧɵɦ ɪɢɫɭɧɤɨɦ ɢ ɬɟɦɧɵɦ ɤɨɪɢɱɧɟɜɵɦ ɩɢɪɚɦɢɞɨɨɛɪɚɡɧɵɦ ɪɢɫɭɧ ɤɨɦ Ƚɨɥɨɜɨɝɪɭɞɶ ɤɨɪɢɱɧɟɜɨ-ɠɟɥɬɚɹ ɫ ɩɪɨɞɨɥɶɧɨɣ ɤɨɪɢɱɧɟɜɨɣ ɩɨɥɨɫɨɣ Ʌɚɩɤɢ ɤɨɪɢɱɧɟɜɵɟ ɫ ɩɨɩɟɪɟɱɧɵɦɢ ɬɟɦɧɵɦɢ ɩɨɥɨɫɤɚɦɢ ɧɚ ɫɨɱɥɟɧɟɧɢɹɯ
ɉɚɭɤɢ A. marmoreus (var pyramidatus) ɪɚɫɩɪɨɫɬɪɚɧɺɧ ɜ ɝɥɭɛɢɧɟ ɫɦɟɲɚɧɧɵɯ ɥɟɫɚɯ ɭɫɬɪɚɢɜɚɟɬ ɥɨɜɱɢɟ ɫɟɬɢ, ɤɚɤ ɧɚ ɥɢɫɬɜɟɧɧɢɱɧɵɯ ɬɚɤ ɢ ɧɚ ɯɜɨɣɧɵɯ ɩɨɪɨɞɚɯ ɞɟɪɟɜɶɟɜ Ƚɨɥɨɜɨɝɪɭɞɶ ɤɨɪɢɱɧɟɜɚɬɨ-ɠɟɥɬɚɹ ɫ ɩɪɨɞɨɥɶɧɨɣ ɤɨɪɢɱɧɟɜɨɣ ɩɨɥɨɫɨɣ ɋɬɪɟɧɭɦ ɤɨɪɢɱɧɟɜɵɣ Ȼɪɸɲɤɨ ɫɜɟɪɯɭ ɫɟɪɨɝɨ ɰɜɟɬɚ ɫ ɧɟɪɟɡɤɨ ɜɵɪɚɠɟɧɧɵɦ ɤɪɟ ɫɬɨɨɛɪɚɡɧɵɦ ɪɢɫɭɧɤɨɦ ɛɟɥɨɝɨ ɰɜɟɬɚ ɜɨ ɜɬɨɪɨɣ ɟɝɨ ɩɨɥɨɜɢɧɟ ɜ ɰɟɧɬɪɟ ɬɟɦɧɨɤɨ ɪɢɱɧɟɜɚɹ ɩɢɪɚɦɢɞɤɚ ɨɤɚɣɦɥɟɧɧɚɹ ɛɟɥɨɣ ɩɨɥɨɫɨɣ
Ⱦɥɹ ɥɨɜɱɢɯ ɫɟɬɟɣ ɩɚɭɤɨɜ A. Marmoreus ɯɚɪɚɤɬɟɪɧɨ ɩɪɚɤɬɢɱɟɫɤɢ ɜɟɪɬɢɤɚɥɶ ɧɨɟ ɪɚɫɩɨɥɨɠɟɧɢɟ ɢɯ ɧɚɞ ɡɟɦɥɟɣ ɭɝɨɥ ɧɚɤɥɨɧɚ ɡɧɚɱɢɬɟɥɶɧɚɹ ɚɫɢɦ ɦɟɬɪɢɹ ɫɟɬɢ ɜɧɢɡ ɢ ɨɬɧɨɫɢɬɟɥɶɧɨ ɤɪɭɩɧɵɟ ɪɚɡɦɟɪɵ ɪɢɫ ɋɟɬɢ ɤɚɤ ɩɪɚɜɢɥɨ ɪɚɫɩɨɥɚɝɚɸɬɫɹ ɧɚ ɫɪɟɞɧɟɨɫɜɟɳɟɧɧɵɯ ɭɱɚɫɬɤɚɯ ɜ ɝɥɭɛɢɧɟ ɥɟɫɚ ɧɚ ɜɵɫɨɬɟ 1±ɦ ɨɬ ɩɨɜɟɪɯɧɨɫɬɢ ɡɟɦɥɢ Ʌɨɜɱɚɹ ɫɟɬɶ A. marmoreus ɹɜɥɹɟɬɫɹ ɤɥɚɫɫɢɱɟɫɤɨɣ ɷɬɚɥɨɧ ɧɨɣ ɜɫɟ ɤɨɦɩɨɧɟɧɬɵ ɫɟɬɢ ɱɟɬɤɨ ɜɵɪɚɠɟɧɵ ɧɚɛɥɸɞɚɟɬɫɹ ɧɢɠɧɹɹ ɩɪɨɫɬɪɚɧɫɬɜɟɧ ɧɚɹ ɚɫɢɦɦɟɬɪɢɹ ɫɟɬɶ ɩɪɟɞɧɚɡɧɚɱɟɧɚ ɞɥɹ ɥɨɜɥɢ ɧɚɫɟɤɨɦɵɯ ɫɪɟɞɧɢɯ ɢ ɦɟɥɤɢɯ ɪɚɡ
ɦɟɪɨɜ ɬɚɛɥ
ɐɟɧɬɪɚɥɶɧɚɹ ɡɨɧɚ ɥɨɜɱɟɣ ɫɟɬɢ ɤɪɭɩɧɚɹ ɫɢɦɦɟɬɪɢɱɧɨɣ ɮɨɪɦɵ ɫ ɧɟɡɧɚɱɢ ɬɟɥɶɧɨɣ ɚɫɢɦɦɟɬɪɢɟɣ ɜɧɢɡ ɫɨɫɬɨɢɬ ɢɡ ɜɢɬɤɨɜ ɱɬɨ ɹɜɥɹɟɬɫɹ ɞɢɚɝɧɨɫɬɢɱɟɫɤɢɦ ɩɪɢɡɧɚɤɨɦ ɜɢɞɚ ɋɜɨɛɨɞɧɚɹ ɡɨɧɚ ɬɚɤɠɟ ɞɨɫɬɚɬɨɱɧɨ ɛɨɥɶɲɚɹ ɩɪɚɜɢɥɶɧɨɣ ɨɜɚɥɶ ɧɨɣ ɮɨɪɦɵ ɫ ɧɟɡɧɚɱɢɬɟɥɶɧɨɣ ɚɫɢɦɦɟɬɪɢɟɣ ɜɧɢɡ ɢ ɜɥɟɜɨ Ʌɨɜɱɚɹ ɡɨɧɚ ɩɥɨɬɧɚɹ ɛɨɥɶɲɚɹ ɩɨ ɪɚɡɦɟɪɚɦ ɫɨ ɡɧɚɱɢɬɟɥɶɧɨɣ ɧɢɠɧɟɣ ɚɫɢɦɦɟɬɪɢɟɣ ɫɨɫɬɨɢɬ ɢɡ ɛɨɥɶɲɨɝɨ ɱɢɫɥɚ ɥɨɜɱɢɯ ɧɢɬɟɣ ɪɚɫɩɨɥɚɝɚɸɳɢɯɫɹ ɧɚ ɪɚɫɫɬɨɹɧɢɢ ɞɪɭɝ ɨɬ ɞɪɭɝɚɫɦ ɱɬɨ ɜɟɪɨɹɬɧɨ ɫɜɹɡɚɧɨ ɫ ɪɚɡɦɟɪɚɦɢ ɧɚɫɟɤɨɦɵɯ ɤɨɦɚɪɵ ɧɚɫɟɤɨ ɦɵɟ ɫɪɟɞɧɢɯ ɪɚɡɦɟɪɨɜ ɫɨɫɬɚɜɥɹɸɳɢɯ ɨɫɧɨɜɭ ɤɨɪɦɨɜɨɣ ɛɚɡɵ ɩɚɭɤɨɜ Ɋɚɡɧɢɰɚ ɜ ɪɚɡɦɟɪɚɯ ɜɟɪɯɧɟɣ ɢ ɧɢɠɧɟɣ ɱɚɫɬɢ ɥɨɜɱɟɣ ɡɨɧɵ ɫɜɹɡɚɧɚ ɫ ɧɚɥɢɱɢɟɦ ɭɛɟɠɢɳɚ ɜ ɜɟɪɯɧɟɣ ɱɚɫɬɢ ɫɟɬɢ ɢ ɩɪɚɤɬɢɱɟɫɤɢ ɜɟɪɬɢɤɚɥɶɧɵɦ ɪɚɫɩɨɥɨɠɟɧɢɟɦ ɫɟɬɢ ɨɬɧɨɫɢ ɬɟɥɶɧɨ ɡɟɦɥɢ ɏɚɪɚɤɬɟɪɧɚɹ ɚɫɢɦɦɟɬɪɢɹ ɩɨɡɜɨɥɹɟɬ ɩɚɭɤɭ ɷɮɮɟɤɬɢɜɧɨ ɞɨɫɬɢɝɚɬɶ ɞɨɛɵɱɭ ɩɨɩɚɜɲɭɸ ɜ ɮɭɧɤɰɢɨɧɚɥɶɧɭɸ ɧɢɠɧɸɸ ɱɚɫɬɶ ɥɨɜɱɟɣ ɡɨɧɵ Ɋɚɞɢɭɫɵ ɜ ɤɨ ɥɢɱɟɫɬɜɟ ɪɚɞɢɚɥɶɧɨ ɪɚɫɯɨɞɹɬɫɹ ɢɡ ɰɟɧɬɪɚ ɢ ɡɧɚɱɢɬɟɥɶɧɨ ɭɞɥɢɧɟɧɵ ɤɧɢɡɭ ȼ ɰɟ ɥɨɦ ɦɨɠɧɨ ɨɬɦɟɬɢɬɶ ɡɧɚɱɢɬɟɥɶɧɵɟ ɪɚɡɦɟɪɵ ɪɚɞɢɭɫɨɜ ɱɬɨ ɜɟɪɨɹɬɧɨ ɫɜɹɡɚɧɨ ɫ ɪɚɫɩɨɥɨɠɟɧɢɟɦ ɫɟɬɟɣ ɧɚ ɞɪɟɜɟɫɧɨɣ ɪɚɫɬɢɬɟɥɶɧɨɫɬɢ
ϯϮ
Ɋɢɫ Ʌɨɜɱɚɹ ɫɟɬɶ A. marmoreus
ϯϯ
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Ɍɚɛɥɢɰɚ |
ɋɪɟɞɧɢɟ ɡɧɚɱɟɧɢɹ ɩɨɤɚɡɚɬɟɥɟɣ ɥɨɜɱɢɯ ɫɟɬɟɣ A. marmoreus |
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ɉɨɤɚɡɚɬɟɥɢ ɫɬɪɭɤɬɭɪɵ ɥɨɜɱɟɣ ɫɟɬɢ |
Ɂɧɚɱɟɧɢɹ ɩɨɤɚɡɚɬɟɥɟɣ ɫɦ |
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ɑɢɫɥɨ ɜɢɬɤɨɜ ɜ ɰɟɧɬɪɚɥɶɧɨɣ ɡɨɧɟ |
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7 |
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Ɋɚɡɦɟɪ ɰɟɧɬɪɚɥɶɧɨɣ ɡɨɧɵ |
ɜɜɟɪɯ |
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ɜɩɪɚɜɨ |
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ɜɧɢɡ |
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ɜɥɟɜɨ |
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Ɋɚɡɦɟɪ ɫɜɨɛɨɞɧɨɣ ɡɨɧɵ |
ɜɜɟɪɯ |
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ɜɩɪɚɜɨ |
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ɜɧɢɡ |
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ɜɥɟɜɨ |
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ɑɢɫɥɨ ɪɚɞɢɭɫɨɜ |
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26 |
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Ⱦɥɢɧɚ ɪɚɞɢɭɫɨɜ |
ɜɜɟɪɯ |
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ɜɩɪɚɜɨ |
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ɜɧɢɡ |
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ɜɥɟɜɨ |
15 |
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Ɋɚɡɦɟɪ ɥɨɜɱɟɣ ɡɨɧɵ |
ɜɜɟɪɯ |
0,60 |
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ɜɩɪɚɜɨ |
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ɜɧɢɡ |
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ɜɥɟɜɨ |
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Ʉɨɥɢɱɟɫɬɜɨ ɥɨɜɱɢɯ ɧɢɬɟɣ |
ɜɜɟɪɯ |
27 |
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ɜɩɪɚɜɨ |
30 |
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ɜɧɢɡ |
39 |
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ɜɥɟɜɨ |
31 |
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Ɋɚɫɫɬɨɹɧɢɟ ɦɟɠɞɭ ɥɨɜɱɢɦɢ ɧɢɬɹɦɢ |
ɜɜɟɪɯ |
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ɜɩɪɚɜɨ |
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ɜɧɢɡ |
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ɜɥɟɜɨ |
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Ɋɚɫɫɬɨɹɧɢɟ ɨɬ ɫɟɪɟɞɢɧɵ ɰɟɧɬɪɚɥɶɧɨɣ ɡɨɧɵ ɥɨɜ |
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ɱɟɣ ɫɟɬɢ ɞɨ ɡɟɦɥɢ |
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ɍɝɨɥ ɧɚɤɥɨɧɚ ɫɟɬɢ ɩɨ ɨɬɧɨɲɟɧɢɸ ɤ ɝɨɪɢɡɨɧ |
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ɬɚɥɶɧɨɣ ɩɨɜɟɪɯɧɨɫɬɢ |
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ɋɬɚɬɢɫɬɢɱɟɫɤɢɣ ɮɚɤɬɨɪɧɵɣ ɚɧɚɥɢɡ ɫ ɢɫɩɨɥɶɡɨɜɚɧɢɟɦ ɦɟɬɨɞɚ ɝɥɚɜɧɵɯ ɤɨɦ ɩɨɧɟɧɬ ɩɨɡɜɨɥɢɥ ɜɵɞɟɥɢɬɶ ɱɟɬɵɪɟ ɤɨɦɩɨɧɟɧɬɵ ɨɩɪɟɞɟɥɹɸɳɢɟ ɫɬɪɭɤɬɭɪɭ ɥɨɜɱɢɯ
ɫɟɬɟɣ A. marmoreus ɬɚɛɥ
ȼɵɞɟɥɟɧɧɵɟ ɤɨɦɩɨɧɟɧɬɵ ɥɨɜɱɢɯ ɫɟɬɟɣ A. marmoreus ɜ ɛɟɪɟɡɨɜɨɦ ɛɢɨɬɨɩɟ ɜɤɥɸɱɚɸɬ ɨɛɳɟɣ ɞɢɫɩɟɪɫɢɢ ɧɟɭɱɬɺɧɧɚɹ ɞɢɫɩɟɪɫɢɹ ɫɨɫɬɚɜɥɹɟɬ
ϯϰ
Ɍɚɛɥɢɰɚ
Ɋɚɫɩɪɟɞɟɥɟɧɢɟ ɨɛɳɟɣ ɞɢɫɩɟɪɫɢɢ ɩɨ ɝɥɚɜɧɵɦ ɤɨɦɩɨɧɟɧɬɚɦ ɜ ɥɨɜɱɢɯ ɫɟɬɹɯ A. marmoreus
Ʉɨɦɩɨɧɟɧɬɵ |
Ɉɛɳɚɹ ɞɢɫɩɟɪɫɢɹ |
1 |
29,5 |
2 |
27,2 |
3 |
9,8 |
4 |
6,4 |
Ɋɟɡɭɥɶɬɚɬɵ ɫɬɚɬɢɫɬɢɱɟɫɤɨɝɨ ɚɧɚɥɢɡɚ ɩɨɥɭɱɟɧɧɵɯ ɪɟɡɭɥɶɬɚɬɨɜ ɩɪɟɞɫɬɚɜɥɟɧɵ ɜ ɬɚɛɥ ȼ ɩɟɪɜɨɣ ɧɚɫɥɟɞɫɬɜɟɧɧɨɣ ɤɨɦɩɨɧɟɧɬɟ ɧɚɛɥɸɞɚɟɬɫɹ ɜɡɚɢɦɨɫɜɹɡɶ ɞɥɢɧ ɪɚɞɢɭɫɨɜ ɢ ɪɚɡɦɟɪɨɜ ɥɨɜɱɟɣ ɡɨɧɵ ɫ ɪɚɫɫɬɨɹɧɢɟɦ ɦɟɠɞɭ ɥɨɜɱɢɦɢ ɧɢɬɹɦɢ ɱɬɨ ɭɤɚ ɡɵɜɚɟɬ ɧɚ ɞɨɦɢɧɢɪɭɸɳɭɸ ɪɨɥɶ ɥɨɜɱɟɣ ɡɨɧɵ ɜ ɫɬɪɭɤɬɭɪɟ ɫɟɬɢ ȼɬɨɪɚɹ ɤɨɦɩɨɧɟɧ ɬɚ ɩɢɳɟɜɚɹ ɨɛɴɟɞɢɧɹɟɬ ɱɢɫɥɨ ɪɚɞɢɭɫɨɜ ɤɨɥɢɱɟɫɬɜɨ ɥɨɜɱɢɯ ɧɢɬɟɣ ɢ ɪɚɫɫɬɨɹɧɢɣ ɦɟɠɞɭ ɥɨɜɱɢɦɢ ɧɢɬɹɦɢ Ⱦɚɧɧɵɣ ɮɚɤɬɨɪ ɨɩɪɟɞɟɥɹɟɬɫɹ ɤɨɥɢɱɟɫɬɜɨɦ ɥɨɜɱɢɯ ɧɢɬɟɣ ɜ ɫɟɤɬɨɪɟ ɢ ɱɢɫɥɨɦ ɪɚɞɢɭɫɨɜ Ɍɪɟɬɢɣ ɮɚɤɬɨɪ ɜɤɥɸɱɚɟɬ ɪɚɡɦɟɪɵ ɰɟɧɬɪɚɥɶɧɨɣ ɢ ɫɜɨɛɨɞɧɨɣ ɡɨɧɵ
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Ɋɚɫɩɪɟɞɟɥɟɧɢɟ ɩɨɤɚɡɚɬɟɥɟɣ ɥɨɜɱɢɯ ɫɟɬɟɣ A. marmoreus |
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Ɍɚɛɥɢɰɚ |
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ɩɨ ɝɥɚɜɧɵɦ ɤɨɦɩɨɧɟɧɬɚɦ |
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ɉɨɤɚɡɚɬɟɥɢ |
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Ɏɚɤɬɨɪ |
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Ɏɚɤɬɨɪ |
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Ɏɚɤɬɨɪ |
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Ɏɚɤɬɨɪ |
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ɑɢɫɥɨ ɜɢɬɤɨɜ ɜ ɰɟɧɬɪɚɥɶ |
|
0,098 |
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0,235 |
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0,401* |
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0,339 |
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ɧɨɣ ɡɨɧɟ |
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Ʉɨɥɢɱɟɫɬɜɨ |
ɜɜɟɪɯ |
|
0,217 |
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0,889* |
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0,177 |
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0,201 |
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ɥɨɜɱɢɯ ɧɢɬɟɣ |
ɜɩɪɚɜɨ |
|
0,131 |
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0,863* |
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0,189 |
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0,059 |
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ɜɧɢɡ |
|
0,308 |
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0,856* |
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0,053 |
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0,016 |
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ɑɢɫɥɨ ɪɚɞɢɭɫɨɜ |
ɜɥɟɜɨ |
|
0,326 |
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0,834* |
|
0,046 |
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0,164 |
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0,144 |
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0,718* |
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0,148 |
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0,192 |
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Ɋɚɡɦɟɪ ɰɟɧ |
ɜɜɟɪɯ |
|
0,475* |
|
0,165 |
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0,559* |
|
0,265 |
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ɬɪɚɥɶɧɨɣ ɡɨɧɵ |
ɜɩɪɚɜɨ |
|
0,381 |
|
0,300 |
|
0,664* |
|
0,238 |
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ɜɧɢɡ |
|
0,401* |
|
0,104 |
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0,657* |
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0,362 |
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ɜɥɟɜɨ |
|
0,371 |
|
0,306 |
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0,654* |
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0,254 |
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Ɋɚɡɦɟɪ ɫɜɨɛɨɞ |
ɜɜɟɪɯ |
|
0,417* |
|
0,441* |
|
0,268 |
|
0,459* |
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ɧɨɣ ɡɨɧɵ |
ɜɩɪɚɜɨ |
|
0,205 |
|
0,372 |
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0,247 |
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0,525* |
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ɜɧɢɡ |
|
0,505* |
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0,529* |
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0,244 |
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0,457* |
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ɜɥɟɜɨ |
|
0,398 |
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0,505* |
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0,313 |
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0,451* |
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ɉɟɪɟɦɟɧɧɚɹ |
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Ɏɚɤɬɨɪ |
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Ɏɚɤɬɨɪ |
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Ɏɚɤɬɨɪ |
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Ɏɚɤɬɨɪ |
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Ɋɚɡɦɟɪ ɥɨɜɱɟɣ |
ɜɜɟɪɯ |
|
0,791* |
|
0,234 |
|
0,274 |
|
0,090 |
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ɡɨɧɵ |
ɜɩɪɚɜɨ |
|
0,755* |
|
0,110 |
|
0,327 |
|
0,079 |
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ɜɧɢɡ |
|
0,898* |
|
0,027 |
|
0,207 |
|
0,091 |
|
|
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|
ɜɥɟɜɨ |
|
0,877* |
|
0,076 |
|
0,200 |
|
0,041 |
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|
Ⱦɥɢɧɚ ɪɚɞɢɭ |
ɜɜɟɪɯ |
|
0,824* |
|
0,240 |
|
0,109 |
|
0,103 |
|
|
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ɫɨɜ |
ɜɩɪɚɜɨ |
|
0,643* |
|
0,340 |
|
0,139 |
|
0,166 |
|
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ɜɧɢɡ |
|
0,809* |
|
0,062 |
|
0,032 |
|
0,134 |
|
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|
ɜɥɟɜɨ |
|
0,731* |
|
0,278 |
|
0,152 |
|
0,102 |
|
|
|
Ɋɚɫɫɬɨɹɧɢɟ |
ɜɜɟɪɯ |
|
0,521* |
|
0,704* |
|
0,094 |
|
0,194 |
|
|
ϯϱ